The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea) Author Shear, William A. Author Richart, Casey H. Author Wong, Victoria L. text Zootaxa 2020 2020-03-19 4753 1 1 78 journal article 21316 10.11646/zootaxa.4753.1.1 f7974bcf-11e7-4d1b-b09d-530dc2dbd54f 1175-5326 3983782 AA9F66B3-EF8C-4F6B-8F35-0BCBEE5122ED Karagama , new genus Type species: Karagama ladybird , n. sp. Diagnosis : Differs from Taiyutyla and the new genus Vancouvereuma in the very simple, acuminate, curved coxites of the posterior gonopods, which lack a T-shaped branch but do have narrow fimbriate regions posteriorly, near a suppressed pore ( Figs. 107, 108 , 110 ). The anterior gonopods ( Figs. 105 , 109 ) are two-branched, similar to those of Bifurcatella , n. gen. (see below) but the branches occupy distinctly anterior and posterior positions, rather than lateral and mesal, as in Bifurcatella species. Etymology : The genus name is an arbitrary combination of letters that should be treated as feminine in gender. Included species : Besides the type , Karagama lupus ( Shear, 2004 ) , n. comb. , and Karagama loftinae ( Shear & Krejca, 2011 ) , n. comb. are also included in the genus. Notes: This genus is somewhat heterogeneous, particularly with regard to the anterior gonopod, which in the type species has the posterior branch tightly appressed to the anterior; while in the other two, it is well separated. Geography may present another problem, since the type species is from northern California ( Redwood National Park ) and the other two included species are from the southern Sierra Nevada and Vancouver Island . Nevertheless for the present, relying only on morphology, they seem to belong together . The name of K. lupus does not change gender since the specific epithet is a noun in apposition rather than an adjective.