New synonyms and redescription of two little-known Thiodia species (Lepidoptera, Tortricidae): T. ursulana (Kennel, 1919) comb. nov. and T. klapperichi (Razowski, 1967) comb. nov.
Author
Bidzilya, Oleksiy
Institute for Evolutionary Ecology of the National Academy of Sciences of Ukraine, 37 Academician Lebediev str., 03143, Kyiv, Ukraine.
Author
Blackstein, Hans
Bukower Weg, 1, D- 14712, Rathenow, Germany.
Author
Budashkin, Yuriy
T. I. Vyazemsky Karadag Scientific Station-Nature Reserve-Branch of FIC “ A. O. Kovalevsky Institute of Biology of the South Seas ” RAS, 298188, Kurortnoye, Pheodosia, Crimea Republic.
Author
Šumpich, Jan
0000-0002-0262-2941
National Museum, Department of Entomology, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jansumpich @ seznam. cz; https: // orcid. org / 0000 - 0002 - 0262 - 2941
jansumpich@seznam.cz
Author
Tsvetkov, Eugene
Mechnikova 3 - 1 - 30, 195271, St. Petersburg, Russia.
text
Zootaxa
2022
2022-04-05
5124
4
458
470
journal article
53640
10.11646/zootaxa.5124.4.3
5b295616-4831-4d3c-8db2-16f0e24233a5
1175-5326
6413668
C88CB391-5086-4680-A06E-1DB457F629A2
Thiodia ursulana
(
Kennel, 1919
)
,
comb. nov.
Figs 1–5
Semasia ursulana
Kennel, 1919: 85
.
Eucosma (Phaneta) ursulana
(
Kennel, 1919
)
—
Obraztsov 1968: 12
.
Eucosma ursulana
(
Kennel, 1919
)
—
Razowski 1999: 475
,
2003: 93
.
ursulana
[no genus, Eucosmini unplaced]—
Gilligan & Wright 2013: 331
.
=
Thiodia aequilibris
Tsvetkov, 2017: 3
,
syn. nov.
Type material.
Lectotype
of
Semasia ursulana
,
♂
, here designated, labelled: “
ursulana Kenn.
Type”, “Uralsk”, “Lectotypus
Semasia ursulana
Kennel, 1919
, des.
O.V. Bidzilya
,
H. Blackstein
,
Yu.I. Budashkin
,
J. Šumpich
,
E.V.
Tsvetkov
, 2022” (
MfN
)
.
Paralectotype
,
♂
, here designated, labelled: “ursulana
n. sp.
Kenn. Type. 89.90.”, “Uralsk”, “
Paralectotypus
Semasia ursulana
Kennel, 1919
, des. O.V. Bidzilya, H. Blackstein, Yu.I. Budashkin, J. Šumpich, E.V.
Tsvetkov, 2022
(
MfN
).
Additional material examined.
♂
,
Indersk
, 28.v, ex coll
Kröne
, gen. slide 18/028, JŠ (
HNHM
)
.
1 ♂
, [without locality data], “
Semasia ursulana Kenn., N. Filipjev
det”, “A. Bang-Haas” (
ZIN
)
.
1 ♂
,
1 ♀
,
Russia
,
Astrakhan Prov.
,
23 km
NE of vill.
Khosheutovo
,
N 47°10’48’’
,
E 48°01’50’’
,
16.v.2018
(
Tsvetkov
) (
RCET
)
;
17 ♂
,
6 ♀
,
Russia
,
Astrakhan Prov.
,
18 km
of vill.
Dosang
,
N 47°00’28’’
,
E 48°06’26’’
,
12.v.2018
(
Tsvetkov
) (
RCET
)
.
The species was originally described in the genus
Semasia
Stephens, 1829
. Later,
Obraztsov (1968:12)
transferred it to the subgenus
Phaneta
Stephens, 1852
of the genus
Eucosma
Hübner, 1823
based on external morphology of adults.
Razowski (1999: 475
;
2003: 93
) followed
Obraztsov (1968)
, but listed
ursulana
under „unplaced species“ of
Eucosma
. He also stated that the species was known only from the original description and that neither the type nor conspecific specimens were found.
Gilligan & Wright (2013: 331)
placed
ursulana
in the group of species with unclear generic assignment (“Eucosmini unplaced”), and noticed that the
holotype
was deposited in MfN (as MNHU).
Semasia ursulana
Kennel, 1919
was described from
two males
collected in the environs of Uralsk (W
Kazakhstan
) (
Kennel 1919: 85
, Pl. III, fig. 17). These
syntypes
, both without abdomens, are kept in the collection of MfN. One of
syntypes
is in good condition, other with both left fore-and hindwings brocken at base. We here designate the better preserved specimen as
lectotype
(see above under “Type material” and
Fig. 1
), and the other (with wings brocken) as
paralectotype
. Another male from Western
Kazakhstan
(Indersk) labelled as “
ursulana
” was found by the authors in the collection of HNHM. The specimen (
Fig. 2
) agrees externally in all details with
two type
specimens of
S. ursulana
, so that we have no doubt of their conspecificity.
Thiodia aequilibris
was described from the male
holotype
collected in
Atyrau Province
and a male
paratype
from
Mangistau Province
of W
Kazakhstan
. The original description is accompanied by the photographs of adults and drawings of the male genitalia (
Tsvetkov 2017: 3–4
, Pl. 1), which fully agree with
S. ursulana
. Hence, the following synonymy is proposed
Thiodia aequilibris
Tsvetkov, 2017
,
syn. nov.
of
Thiodia ursulana
(
Kennel, 1919
)
,
comb. nov.
Remarks.
The assignment of
ursulana
to the genus
Thiodia
and the position of the species within the genus was discussed by
Tsvetkov (2017: 4
, as
T. aequilibris
). The species can be clearly separated from its congeners both by the male genitalia (
Fig. 4
) and by the wing pattern. The female of
T. ursulana
was illustrated in the original description of
T. aequilibris
(
Tsvetkov 2017: 4
, Pl. 1, fig. 2). However, the specimen was later lost, but its genitalia have been described, though not illustrated. Below we provide a drawing and re-description of the female genitalia of
T. ursulana
based on additional material from
Astrakhan Province
of
Russia
.
Female genitalia
(
Fig. 5
). Papillae anales elongate, covered by small bristles. Posterior apophyses nearly equal to anterior apophyses, strongly broadened in posterior 1/2. Lamella postvaginalis short and small, anterior margin with a tiny, sharp central protrusion, posterior margin straight. Lamella antevaginalis small, evenly bent or V-shaped. Antrum small, heart-shaped. Sternum VII trapezoidal with bent sides and rounded angles, two vane-like protrusions of posterior margin vary in length. Ductus bursae membranous, cingulum tapering posteriorly, weakly sclerotised or membranous ventrally, usually obliquely cut anteriorly, covered by sternum VII (in ventral view). Corpus bursa spherical, membranous, with a small horn-like signa.
FIGURES 1–5.
Thiodia ursulana
(
Kennel, 1919
)
. 1. Lectotype. Uralsk. 2. Male, Indersk (gen. slide 18/028, JŠ). 3. Female, Russia, Astrakhan Prov., 18 km of vill. Dosang, N 47°00’28’’, E 48°06’26’’, 12.V.2018. 4. Male genitalia, Indersk (gen. slide 18/028, JŠ). 5. Female genitalia, Russia, Astrakhan Prov.
Distribution.
Kazakhstan
: Western
Kazakhstan
,
Atyrau
and Mangystau Provinces;
Russia
:
Astrakhan Province
(new record),
Buryatia
: Munku-Sardyk Mountain (
Filipjev 1929: 6
).