The evolution of hermit crabs (Crustacea, Decapoda, Anomura, Paguroidea) on the basis of carapace morphology: a state-of-the-art-report Author Fraaije, René H. B. Author Van Bakel, Barry W. M. Author Jagt, John W. M. Author Charbonnier, Sylvain Author Schweigert, Guenter Author Garcia, Géraldine Author Valentin, Xavier text Geodiversitas 2022 2022-01-06 44 1 1 16 journal article 2657 10.5252/geodiversitas2022v44a1 2d127866-508b-4d25-9d03-0db12cfe11be 1638-9395 5834191 urn:lsid:zoobank.org:pub:6A4424D0-DEFD-4783-9B75-24AC2C7829F7 Family PROBEEBEIDAE Boone, 1926 , emended herein Probeebeidae Boone, 1926b: 73 . TYPE GENUS. — Probeebei (Boone, 1926) by monotypy. OTHER GENERA INCLUDED. — Labidochirus Benedict, 1892 , Tisea Morgan & Forest, 1991 and Tylaspis Henderson, 1885 . DIAGNOSIS. — Shield (excluding rostrum) strongly convex, well calcified, width equal to length or width exceeding length, with distinct bulges (i.e., keraial and massetic regions) laterally, strong to weaker spinose ornament. Rostrum well developed, exceeding lateral projections. Cervical and branchial grooves subparallel, encompassing small branchial regions. Posterolateral margins spinose. Posterior carapace well calcified, broadly inflated, with dense spinose ornamentation. Well-developed cardiac grooves encompassing cardiac region. REMARKS. — The bulges on the shield of extant probeebeids ( Fig. 10 ) are quite similar to those of the most basal paguropsid, Eopaguropsis nidiaquilae (see Fraaije et al. 2012c : figs 2c; 3), which also has a spinose ornamentation on the gastric and lateral parts of the shield.