Impatiens bokorensis (Balsaminaceae), a new species from Cambodia Author Cho, Seong-Hyun International Biological Material Research Center, Korea Research Institute of Bioscience and Biotechnology, 125 Gwahak-ro, Yuseong-gu, Daejeon 34141, Republic of Korea Author Kim, Bo-Yun Department of Life Science, Hallym University, 1 Hallymdaehak-gil, Chuncheon-si, Gangwon 24252, Republic of Korea Author Park, Han-Sol Department of Life Science, Hallym University, 1 Hallymdaehak-gil, Chuncheon-si, Gangwon 24252, Republic of Korea Author Phourin, Chhang Forestry Administration, 40 Preah Norodom Blvd, Phnom Penh, Kingdom of Cambodia Author Kim, Young-Dong https://orcid.org/0000-0001-5904-4727 Department of Life Science, Hallym University, 1 Hallymdaehak-gil, Chuncheon-si, Gangwon 24252, Republic of Korea ydkim@hallym.ac.kr text PhytoKeys 2017 2017-02-07 77 33 39 http://dx.doi.org/10.3897/phytokeys.77.11345 journal article http://dx.doi.org/10.3897/phytokeys.77.11345 1314-2003-77-33 FFA6430F842AFFFCFFA8FFF6FFFE6013 269674 Impatiens bokorensis S.H.Cho & B.Y.Kim sp. nov. Figures 1 , 2 Type . CAMBODIA . Kampot Province , Phnum Bokor National Park , sandstone tables in evergreen forest margin, 10°38'20.8"N , 104°00'16.0"E , a.s.l. 1,050 m , 24 August 2015 , with flowers, Cho S.H , Kim B.Y. , Park H.S. , Chhang Phourin CB-3112 ( holotype HHU!, isotypes KB!, KRIB!, RUPP!) . Diagnosis . Impatiens bokorensis is most similar to the Thailand endemic species Impatiens patula Craib in habit but is readily distinguished from the latter by the orbicular-obovate dorsal petal, shorter pedicels and larger seeds (Table 1 ). Table 1. Comparison of key features of Impatiens bokorensis and Impatiens patula .

Taxonomic traits	Impatiens bokorensis	Impatiens patula
Leaf	-	-
upper surface	pubescent	pubescent with scurfy hairs
lower surface	glabrous to sparsely pubescent	pubescent on nerves only or tomentose all over
strigose-ciliate at base	1-2.7 mm	1-1.4 mm
Pedicel	1.4-2.0 cm	2.3-3(-5) cm
Dorsal petal	orbicular-obovate, ca. 6 x 8 mm, horned at apex, horn 2.2-2.3 mm long	cordate, 7 mm long, horned at apex, horn 3 mm long
Seeds	3.8-4.6 x 2.6-3.2 mm	3.5 x 2.5 mm

Description. Herbs, annual, terrestrial, hermaphroditic. Stems erect, 15-40 cm tall, tinged purplish red, branched, glabrous or sparsely puberulous on the upper part. Leaves simple, alternate; petioles subsessile to 1.4 mm; leaf blade lanceolate to ovate-lanceolate, apex acuminate, base narrowly cuneate to attenuate, 2.5-7.0 x 0.6-2.0 cm, upper surface pubescent, lower surface glabrous to sparsely pubescent, secondary veins pinnate, 6 to 8 on each side of mid-vein, margin serrate, teeth mucronate and purple tinged, strigose-ciliate at base; strigose-ciliate, 4-14, 1-2.7 mm long, purple to purplish black, minutely puberulous at base. Flowers axillary, solitary, rarely 2 fascicled, zygomorphic, minutely puberulous; pedicels slender, erect, purplish red, 1.4-2.0 cm long, glabrous, bracteate at base; bracts linear, up to 4 mm; lateral sepals 2, linear-lanceolate, 2.5-3 mm long, glabrous; lower sepal funnel-form, pink, ca. 5 mm long, ca. 3 mm deep; spur 17-23 mm long, slightly curved; dorsal petal, orbicular-obovate, ca. 6 x 8 mm, horned at apex, horn 2.2-2.3 mm long; lateral united petals separate, bilobed, ca. 11 mm long; upper petals oblong, 6.0-6.5 x ca. 3.0 mm, minutely apiculate; lower petals, 9.0-9.3 x 3.5-3.8 mm; androecium ca. 2.8 x 1.6 mm; stamens 5, connate, surrounding gynoecium; filaments ca. 0.7 mm; ovary fusiform, pubescent, ca. 2.5 x 1.0 mm; style glabrous, ca. 0.2 mm long; stigma 5, ca. 0.25 mm long. Fruit a capsule, fusiform, ca. 15 x 6 mm, pubescent with scurfy hairs, 3[4]-seeded. Seeds obovoid, slightly compressed, 3.8-4.6 x 2.6-3.2 mm, pubescent with spirally sculptured hairs. Specimen examined. CAMBODIA. 16 November 2015, with fruits, Cho et al. CB-3432 (HHU!, KRIB!); 2 September 2016, with flowers, Kim et al. CB-3537 (HHU!) Phenology. Flowering specimens were collected in August and fruiting specimens in November. Distribution and habitat. Impatiens bokorensis grows on sandstone tables in evergreen forest margins at 1,050 m a.s.l.. Endemic to southwestern Cambodia, Impatiens bokorensis is at present known only in the type locality. GenBank Accession No. Cho et al. CB-3432: KX171761 (ITS). Conservation status. Impatiens bokorensis was collected in Phnum Bokor National Park in southwestern Cambodia. Until now, only one population, consisting of ca. 200 individuals, has been discovered in the park area; therefore, it is preliminarily classified as data deficient (DD) according to the IUCN Red List criteria (IUCN 2001).