Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica Author Fleming, AJ Author Wood, D. Monty Author Smith, M. Alex Author Hallwachs, Winnie Author Janzen, Daniel H text Biodiversity Data Journal 2018 6 11223 11223 http://dx.doi.org/10.3897/BDJ.6.e11223 journal article http://dx.doi.org/10.3897/BDJ.6.e11223 1314-2828-6-11223 Calolydella erasmocoronadoi Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039246 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039246; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20642, BOLD:AAC8991, ASTAV809-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 14-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011706 ; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza ; individualID: DHJPAR0011706; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-2399, BOLD:AAC8991, ASTAS432-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Huerta; verbatimElevation: 527 ; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 17-Jun-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037554 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0037554; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-69806, BOLD:AAC8991, ASHYC4299-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 10-Sep-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011688 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011688; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55482, BOLD:AAC8991, ASTAS414-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 19-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039244 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039244; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20643, BOLD:AAC8991, ASTAV807-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039242 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039242; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20622,, ASTAV805-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039245 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039245; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20618, BOLD:AAC8991, ASTAV808-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037556 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0037556; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-69793, BOLD:AAC8991, ASHYC4301-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 10-Sep-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037555 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0037555; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-69805, BOLD:AAC8991, ASHYC4300-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 10-Sep-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039243 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039243; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20621, BOLD:AAC8991, ASTAV806-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011686 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011686; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55195, BOLD:AAC8991, ASTAS412-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 02-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011687 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011687; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55483, BOLD:AAC8991, ASTAS413-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 20-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011685 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011685; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55194, BOLD:AAC8991, ASTAS411-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 03-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039251 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039251; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20639, BOLD:AAC8991, ASTAV814-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039250 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039250; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20619, BOLD:AAC8991, ASTAV813-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011707 ; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza ; individualID: DHJPAR0011707; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-2393, BOLD:AAC8991, ASTAS433-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Huerta; verbatimElevation: 527 ; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 03-Jun-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011689 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011689; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55450, BOLD:AAC8991, ASTAS415-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 18-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039248 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039248; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20624, BOLD:AAC8991, ASTAV811-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037553 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0037553; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-69808, BOLD:AAC8991, ASHYC4298-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 08-Sep-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011708 ; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza ; individualID: DHJPAR0011708; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-2401, BOLD:AAC8991, ASTAS434-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Huerta; verbatimElevation: 527 ; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 17-Jun-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039247 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039247; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20640, BOLD:AAC8991, ASTAV810-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Monte Cristo; verbatimElevation: 525 ; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 14-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039249 ; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios ; individualID: DHJPAR0039249; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-20623, BOLD:AAC8991, ASTAV812-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste ; verbatimLocality: Quebrada Serrano; verbatimElevation: 585 ; verbatimLatitude: 11.0002; verbatimLongitude: -85.4561; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 11-May-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037558 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0037558; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-69807, BOLD:AAC8991, ASHYC4303-10; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 14-Sep-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011684 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: DHJPAR0011684; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55193, BOLD:AAC8991, ASTAS410-06; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 04-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 04-SRNP-55202 ; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios ; individualID: 04-SRNP-55202; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 04-SRNP-55202; Taxon: scientificName: Calolydellaerasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Mismo; verbatimElevation: 680 ; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe ; verbatimEventDate: 04-Oct-2004 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Male (Fig. 15a, b, c). Length: 6-8mm. Head (Fig. 15b): frontal setae extending to base of postpedicel; fronto-orbital plate up to 50% silver pollinose, almost bare, with a few small setulae interspersed among frontal setae; parafacial silver throughout. Thorax (Fig. 15a, c): gold on dorsal surface, silver laterally (>50% coverage); thoracic vittae visible only under certain angles of light; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 1:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6-7 setulae dorsally, extending halfway to crossvein R-M. Abdomen (Fig. 15a): ground color black with uninterrupted transverse gold pollinose bands; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 16): sternite 5 (Fig. 16c) with two small lobes and a wide U-shaped median cleft, 0.34X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; upper half of sternite with many stout, outwardly-pointing setae. Cerci (Fig. 16b), in dorsal view, separated by a narrow gap; each cercus long and rounded, and not significantly tapering from base to tip; cercus straight when viewed laterally, with a slight downward hook at its tip, setose almost along its entire length. Surstylus (Fig. 16c) stout and digitiform; apical half with short setae; tip of surstylus not lobed or angled inwards when viewed dorsally; subequal in length to cercus. Female (Fig. 15d, e, f). Length: 4-7mm. Fronto-orbital plate 2.5X as wide as in male. Diagnosis Calolydella erasmocoronadoi can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate bare, up to 50% silver pollinose, thoracic vittae visible only under certain angles of light, anatergite with three or more setae arranged in a small tuft, wing vein R4+5 with 6-7 setulae, and abdomen with uninterrupted transverse marginal pollinose bands. Etymology The specific epithet is in honor of Erasmo Coronado Caballo of Liberia, Costa Rica, in recognition of his high-quality assistantship to Sigifredo Marin of Liberia and to the ACG parataxonomists who collected and reared the caterpillars from which the new species of ACG Calolydella described here emerged. Distribution Costa Rica, ACG, Guanacaste Province, Sendero Mismo, 680m. Ecology Calolydella erasmocoronadoi has been reared 42 times from Euselasia chryssipe (Bates, 1866) ( Lepidoptera : Riodinidae ), in rain forest.