Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae) Author Frolov, Andrey V. Author Akhmetova, Lilia A. Author Scholtz, Clarke H. text Journal of Natural History 2008 2008-06-30 42 21 - 24 1477 1508 http://dx.doi.org/10.1080/00222930802002535 journal article 10.1080/00222930802002535 1464-5262 5219397 Coptorhina nitefacta Gillet ( Figures 5 , 13H , 22 ) Coptorhina nitefacta Gillet 1932:327 ; Ferreira 1972:357 ; 1939:34, 38. Type material examined Coptorhina nitefacta . Holotype „ with labels ‘‘AFR. OR. ANGL. (WA-TaïTA) BURA Alluaud and Jeannel Mars 1912 – 1050 m – St. 61’’, ‘‘ Coptorhina nitefacta Gillet type.’’, ‘‘Holotype’’ ( MNHN ) . Additional material examined Kenya : Garissa env. [00 ° 279 S 39 ° 399 E], 30 November 1999 , M. Snižek leg. 1 spm. ( JJCM ) . Diagnosis This species differs from other similar Coptorhina species in having shiny, very sparsely punctate upper side of body. Description Strongly convex, black, shiny beetle. Body length 9.5–13.5 mm, width 6.5–9.5 mm. Dorsal surface without visible setae. Head . Clypeus with protruding anterior angles separated by deep, right-angled sinuation ( Figure 5 ). Protruding angles acute to narrowly rounded apically, with longitudinal carinae on the upper side. Genae obtusely rounded. Genal sutures feebly distinct. Lateral margins of clypeus sinuate near genal sutures in some individuals. Frontal suture feebly visible, broadly interrupted in the middle. Head very densely and evenly punctuate, punctures sometimes adjacent and sculpture appears rugose. Antennal clubs dark-brown. Pronotum . Trapezoidal, two times wider than long. Anterior margin and base finely bordered. Sides more or less evenly rounded. Lateral border relatively narrow, evenly tapering basally and apically. Pronotum divided into larger anterior part and smaller posterior part by a transversal carina. The carina is almost as wide as pronotum, narrowly interrupted medially, not serrate. Anterior part of pronotum punctate, excavated laterally in larger specimens. Posterior part sparsely punctate with minute punctures. Elytra . Striae fine but distinct, punctate (punctures separated by 3–4 puncture diameters). Intervals flat, very sparsely punctate with minute punctures. Underside . Pygidium with fine border, its disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of metasternum which is very sparsely to almost indistinctly punctuate. Aedeagus . Parameres with widely rounded apices in lateral view. Variability The description above is based mostly on the holotype . The other specimen known to us is smaller and has less developed transversal pronotal carina and slightly or not excavated anterolateral sides of pronotum. It is however impossible to appreciate the interspecific variability from the material available. Distribution This species is known from two localities in Kenya ( Figure 22 ).