<p> <strong> A revision of Afrotropical Quasimodo flies (Diptera: Schizophora; Curtonotidae). Part IV — the continental Afrotropical species of <em> Curtonotum </ em> Macquart, with descriptions of thirteen new species and a combined phylogenetic analysis of the Curtonotidae </ strong> </ p> Author Kirk-Spriggs, Ashley H. ashley.kirk-spriggs@nasmus.co.za Author Wiegmann, Brian M. ashley.kirk-spriggs@nasmus.co.za text Zootaxa 2013 2013-07-09 3684 1 1 166 http://dx.doi.org/10.11646/zootaxa.3684.1.1 journal article 10.11646/zootaxa.3684.1.1 1175-5326 5298914 E922034E-1247-400B-97F6-1778CF766B91 Curtonotum cimbebas Kirk-Spriggs , sp. n. Figs 123 , 147 , 183 , 269, 272, 275 , 331 . Curtonotum cuthbertsoni : Duda, 1935: 31 , Type VII of Tsacas (1977: 153) . Etymology . The specific epithet cimbebas is a noun in apposition, derived from “Cimbebas”; the archaic name for the area between Cape Negro and the Tropic of Capricorn on the southwest African seaboard. Description : Male (primarily based on field-pinned HT). As described for C . uncinatum (above), differing in the following respects: Measurements : Overall length unknown; length of head and thorax combined 2–2.7 mm ; length of thorax and scutellum combined 1.9–2.6 mm ; wing length 3.3–3.8 mm ( n = 5, HT, PT). Head ( Figs 123 , 147 ). Eye height/length ratio: 11:6 ( n = 1, HT); frons ( Fig. 147 ) wider than long, frons length/ width ratio: 6:8 ( n = 1, HT), very slightly wider at vertex than at ventral margin; arista with 7–9 long dorsal branches and 2 or 3 ventral branches; gena narrow eye height/genal height ratio: 11:1 ( n = 1, HT). Thorax ( Fig. 123 ). Anepisternum with 13 fine setulae 3 arranged in a group in posterior corner; katepisternum with 14 short, fine setulae. Legs . Fore coxa with 7 brown setulae; fore tibia with ctenidium of 9–10 short, sharp, black spinules. Wing ( Fig. 183 ). Dm–cu crossvein vertical with even arch; anal lobe broad. Abdomen . Sternite 6 as in Fig. 305 . Terminalia ( Figs 269, 272, 275 ). Hypandrium ( Fig. 269 , hy ); postgonite ( pg ); epandrium ( ep ); cercus ( ce ); surstylus ( ss ); phallus (as in Figs 272 , ph , bp , dp ; 275, bp , dp ); phallapodeme ( Fig. 272 , ph ); ejaculatory apodeme ( ea ); basiphallus ( Figs 272, 275 , bp ) wide basally, expanded into quite broad lateral expansion apically; distiphallus ( dp ) long, scimitar-like (viewed laterally), membranous area broad, basal “spine” very short, but erect and sharp, more apical spine relatively broad basally and sharp. ♀ Unknown. Variation . No significant variation is noted. Differential diagnosis . Curtonotum cimbebas sp. n. belongs to a group of ten species here ascribed to the uncinatum species-group, nine of which are virtually identical externally and are only separable based on minor differences in the male phallus (see Differential diagnosis under C . uncinatum sp. n. for details). Curtonotum cimbebas sp. n. can be separated from other species in the species-group by reference to the above key. Type material examined . NAMIBIA : holotype ♂ , “ Namibia : KARIBIB DISTRICT. / Tsaobismund 85 at: / 22°22'40"S 15°44'58"E / 13-15.iv.2001 / A.H. Kirk-Spriggs & E. Marais / Malaise traps // Namibian National / Insect Collection / National Museum / P.O. Box 1203 / Windhoek, Namibia [printed; blue card] // HOLOTYPE ♂ / Curtonotum / cimbebas sp. n. / A.H. Kirk-Spriggs 2011 [printed; red card]” (NMNW). In excellent condition; dissected, abdomen and terminalia in micro-vial pinned beneath specimen. Paratypes (all labelled: “ PARATYPE / Curtonotum ♂ / cimbebas sp. n. / A.H. Kirk-Spriggs 2011 [printed; blue card]”): NAMIBIA : 9♂ , same data as holotype ; 1♂ , “ Namibia : NAUKLUFT PK / Tsams Ost spring / 24°14'45"S , 16°06'17"E / 26-27.xi.1997 / [A.H.] Kirk-Spriggs & [E.]. Marais / Malaise trap”; 5♂ , “ Namibia : NAUKLUFT PK / Naukluft spring / 24°15'78"S, 16°14'08"E / 28-30.xi.1997 / [A.H.] Kirk-Spriggs & [E.] Marais / Malaise trap”; 5♂ , “ Namibia : WINDHOEK DISTRICT / Greylingshof 107: Gaub Canyon / 23°29'S 16°46'E / 02-03.x.1999 / A.H. Kirk-Spriggs & T. Pape / Malaise trap sample” (all NMNW); 5♂ , “South West Africa / Gobabeb 30.I.1978 / Kuiseb River Bed / O. Lomholdt leg.” (ZMUC). Distribution . Endemic to Namibia ( Fig. 331 ). Apparently confined to the Namibian Escarpment and immediate vicinity. Bionomics . Occurring in the Nama Karoo and Namib Desert major habitat types ; predominately in the Deserts and Xeric Shrublands vegetation type (Appendix III ). Sampled in Malaise traps in the vicinity of springs in the Naukluft Mountains , which form part of the Namibian Escarpment , and in the ephemeral riverbeds of the Kuiseb and Tsaobis Rivers of Namibia .