Revision of the western Palaearctic species of Aleiodes Wesmael (Hymenoptera, Braconidae, Rogadinae). Part 2: Revision of the A. apicalis group
Author
van Achterberg, Cornelis
Author
Shaw, Mark R.
Author
Quicke, Donald L. J.
text
ZooKeys
2020
919
1
259
http://dx.doi.org/10.3897/zookeys.919.39642
journal article
http://dx.doi.org/10.3897/zookeys.919.39642
1313-2970-919-1
0CC5169A232541AD938F179FCB056381
CBA7303BD1B65E07A4DC591C877727BA
Aleiodes Wesmael, 1838
Figs 10-21
, 22-25
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Aleiodes
Wesmael, 1838: 194;
Shenefelt 1975
: 1163-1185;
Marsh 1979
: 177-178;
Papp 1985a
: 143-164,
1985b
: 347-349;
Shaw and Huddleston 1991
: 95-96 (biology);
van Achterberg 1991
: 24;
Zaldivar-Riveron
et al. 2004
: 225,
2008
: 329 (phylogeny);
van Achterberg and Shaw 2016
: 8-11 (groups). Type species (designated by Viereck 1914):
Aleiodes heterogaster
Wesmael, 1838 [examined; =
A. albitibia
(
Herrich-Schaeffer
, 1838)].
Petalodes
Wesmael, 1838: 123;
Tobias 1971
: 218 (transl. 1975: 86-87);
Shenefelt 1975
: 1209-1211;
Tobias 1976
: 90;
Marsh 1979
: 179;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838);
van Achterberg and Shaw 2016
: 8. Type species (by monotypy):
Petalodes unicolor
Wesmael, 1838 [examined; =
Aleiodes compressor
(
Herrich-Schaeffer
, 1838)].
Schizoides
Wesmael, 1838: 94. Unavailable name.
Nebartha
Walker, 1860: 310;
Shenefelt 1975
: 1216;
Marsh 1979
: 179;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Nebartha macropodides
Walker, 1860 [examined].
Tetrasphaeropyx
Ashmead, 1889: 634;
Shenefelt 1975
: 1260;
Marsh 1979
: 179;
Fortier and Sherman 2008
: 445 (as subgenus of
Aleiodes
Wesmael, 1838);
Zaldivar-Riveron
et al. 2008
: 329 (as synonym of subgenus
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Rogas pilosus
Cresson, 1872 [examined].
Neorhogas
Szepligeti
, 1906: 605;
Shenefelt 1975
: 1205;
van Achterberg 1991
: 24 (as subgenus of
Aleiodes
Wesmael, 1838);
Zaldivar-Riveron
et al. 2008
: 329 (included in subgenus
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Neorhogas luteus
Szepligeti
, 1906 [examined; =
Aleiodes praetor
(Reinhard, 1863)].
Chelonorhogas
Enderlein, [Sept. 1st] 1912a: 258;
Shenefelt 1975
: 1187;
van Achterberg 1991
: 24 (as subgenus of
Aleiodes
Wesmael, 1838);
Zaldivar-Riveron
et al. 2008
: 329 (as subgenus of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Chelonorhogas rufithorax
Enderlein, 1912 [examined; not
Aleiodes rufithorax
(Cameron, 1911) =
A. convexus
van Achterberg, 1991].
Eucystomastax
Brues, [(end of?) Sept.] 1912: 223;
Shaw 1993
: 5 (as subgenus of
Aleiodes
Wesmael, 1838);
Zaldivar-Riveron
et al. 2004
: 225 (included in
Aleiodes
Wesmael, 1838); Shimbori & Penteado-Dias 2011: 17 (as subgenus of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Eucystomastax bicolor
Brues, 1912 (=
Rogas melanopterus
Erichson, 1848).
Leluthinus
Enderlein, 1912b: 96;
Shenefelt 1975
: 1202-1203;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Leluthinus lividus
Enderlein, 1912 [examined].
Aleirhogas
Baker, 1917b: 383, 411;
Shenefelt 1975
: 1185-1186;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838). Type species (designated by Viereck 1921):
Rhogas (Aleirhogas) schultzei
Baker, 1917 [examined].
Hemigyroneuron
Baker, 1917a: 284, 322-327;
Zaldivar-Riveron
et al. 2008
: 329 (as subgenus of
Aleiodes
Wesmael, 1838);
Butcher and Quicke 2011
: 1405 (as subgenus of
Aleiodes
Wesmael, 1838, and
Hemigyroneuron
sensu
Zaldivar-Riveron
et al. (2008)
is not
Hemigyroneuron
));
Butcher and Quicke 2015
: 275-279. Type species (original designation):
Hemigyroneuron speciosus
Baker, 1917 [examined].
Heterogamoides
Fullaway, 1919: 43;
Shenefelt 1975
: 1188;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Heterogamoides muirii
Fullaway, 1919 [examined].
Cordylorhogas
Enderlein, 1920: 153;
Shenefelt 1975
: 1195;
van Achterberg 1991
: 31;
Zaldivar-Riveron
et al. 2004
: 232,
2008
: 329 (as synonym of subgenus
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Cordylorhogas trifasciatus
Enderlein, 1920 [examined].
Hyperstemma
Shestakov, 1940: 10;
Shenefelt 1975
: 1200;
van Achterberg 1991
: 24 (as synonym of
Aleiodes
Wesmael, 1838). Type species (by monotypy):
Hyperstemma chlorotica
Shestakov, 1940 [examined].
Dimorphomastax
Shenefelt, 1979: 131-133;
Shaw et al. 1998
: 66 (as synonym of
Aleiodes
Wesmael, 1838). Type species (by original designation):
Dimorphomastax peculiaris
Shenefelt, 1979 [examined; =
Aleiodes atriceps
Cresson, 1869].
Pholichora
van Achterberg, 1991: 48-53;
Quicke and Shaw 2005
: 532;
Zaldivar-Riveron
et al. 2008
: 329 (as synonym of
Aleiodes
Wesmael, 1838);
Butcher and Quicke 2011
: 1405 (as synonym of subgenus
Hemigyroneuron
Baker, 1917);
Butcher et al. 2012
: 9 (id.). Type species (original designation):
Hemigyroneuron madagascariensis
Granger, 1949 [examined].
Arcaleiodes
Chen & He, 1997: 60-62;
Zaldivar-Riveron
et al. 2008
: 329 (as subgenus of
Aleiodes
Wesmael, 1838);
Butcher et al. 2012
: 18-19 (id.). Type species (original designation):
Aleiodes unifasciatus
Chen & He, 1991 [examined].
Vietorogas
Long & van Achterberg, 2008: 313-314;
Butcher et al. 2012
: 15-17 (as synonym of
Aleiodes
Wesmael, 1838). Type species (original designation):
Vietorogas bachma
Long, 2008 [examined].
R
(
h
)
ogas
auct; Tobias, 1971: 215-217 (transl. 1975: 83-86); Shenefelt, 1975: 1215-1256; Tobias, 1976: 81-89; Marsh, 1979: 179-181; Tobias, 1986: 74-84.
Notes.
Hyperstemma
Shestakov, 1940, is traditionally included in the genus
Heterogamus
Wesmael, 1838 (
Shenefelt 1975
) or in the subgenus
Heterogamus
of the genus
Aleiodes
Wesmael (e.g.,
Belokobylskij 2000
), but differs by the shape of the head (Figs
17-19
) and of the tarsal claws (Fig.
20
), the position of the clypeus (Fig.
17
), the elongate 2nd submarginal cell of the fore wing (but folded in Fig.
10
), the widened 1st subdiscal cell of the fore wing and distinctly widened marginal cell of the hind wing (Fig.
10
). Therefore, we retain the subgenus
Hyperstemma
Shestakov of
Aleiodes
Wesmael for at least the following species:
A. chloroticus
(Shestakov, 1940) from China (Palaearctic and Oriental), *Japan (RMNH), Korea, and Russia (Far East),
A. albigenus
Chen & He, 1997, from China (Oriental) and Vietnam,
A. crassinervis
Chen & He, 1997, from China (Oriental) and Vietnam,
A. naevius
Chen & He, 1997, from China (Oriental), and
A. pallidinervis
(Cameron, 1910) from China (Palaearctic and Oriental), Japan, Korea, and Russia (Far East).
Figures 10-21.
Aleiodes chloroticus
(Shestakov), ♂, Japan
10
wings
11
mesosoma lateral
12
mesosoma dorsal
13
1st-3rd metasomal tergites dorsal
14
fore femur lateral
15
hind femur lateral
16
base of antenna
17
head anterior
18
head dorsal
19
head lateral
20
outer hind tarsal claw
21
apex of antenna.