The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae Author Beaver, Roger A. 161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand. Author Smith, Sarah M. 0000-0002-5173-3736 Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736 smith462@msu.edu text Zootaxa 2022 2022-08-08 5174 1 1 24 journal article 115950 10.11646/zootaxa.5174.1.1 c4bf1061-5af8-4f4a-b4ca-234b6d0ca615 1175-5326 6972960 F80F144B-D1E8-4587-A146-0BACFFE18FB6 Polygraphus major Stebbing Polygraphus major Stebbing, 1903: 234 . Recorded from Bhutan by Wood & Bright (1992) . We have been unable to trace the original record, but it is highly probable that the species occurs in the country. Distribution. Bhutan , China ( Xizang ), India ( Himachal Pradesh , Punjab, Uttarakhand ), Kashmir, Nepal , Pakistan . Biology. This species attacks conifers in the genera, Pinus , Picea and Cedrus (Pinaceae) , but is particularly important as a pest of Pinus wallichiana in the Himalayan region ( Browne 1968 ). Its biology has been described by Stebbing (1914) , Beeson (1961) and Khanday et al. (2020) . Associated coleopteran predators are considered to be important in regulating population levels ( Khanday et al. 2016 , 2018 ).