First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group Author Kamiński, Marcin J. Author Gearner, Olivia M. Author Kanda, Kojun Author Swichtenberg, Kali Author Purchart, Luboš Author Smith, Aaron D. text Zoological Journal of the Linnean Society 2021 2020-06-16 191 883 901 journal article 3320 10.1093/zoolinnean/zlaa052 fd4b2931-7bad-4644-b485-f365040ae01e 0024-4082 5724192 0810A91E-7D8D-4F56-B468-9D5B3A6115E9 KEY TO THE SPECIES OF THE GENUS TOKTOKKUS 1. Apex of elytra not sloped (dorsal view) and depressed with clear margin ( Fig. 3B )............... T. congolensis - Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g. Fig. 3A ) ...............................................2 2. Margin of prosternal collar expanded and folded out into a large lip ( Fig. 3C ); elytra round, almost heart- shaped ( Fig. 3C ); declivous portion of elytra with little to no tubercles..................................... T. tschinkeli - Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows .........................................................................................................3 3. Disc of pronotum with prominent punctures ( Fig. 3D ); gold setae present on elytra ( Fig. 3D )..... T. herero - Disc of pronotum only with micropunctures; elytra not covered with setae................................................4 4. Tubercles on elytral sides round/globular, vertically direct ( Fig. 2B, C ) ......................................................5 - Tubercles on elytral sides pointed, directed posteriorly (e.g. Fig. 3F ) ..........................................................6 5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round ( Fig. 2B ) ........................................................................................................................................................... T. mulleri - Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round ( Fig. 2C )................................................................................................................................ T. schultzei 6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7 - Elytral tubercles dense (more than 6 lateral rows), slightly angled ( Fig. 2B–F ).........................................8 7. Elytral tubercles distinct, not confluent, nearly reaching humeri ( Fig. 3A ) ................................ T. barclayi - Elytral tubercles small and short, confluent into rows, terminating well before humeri ( Fig. 2A ) ...................................................................................................................................................... T. mashunus 8. Microtubercles present between tuberculate rows (e.g. Fig. 3E ), tuberculate rows rarely elevated on ridges ...............................................................................................................................................................9 - Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges ................................................................................................................................10 9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short .................................................................................................................... T. tuberculipennis - Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller ................................................................................................................................ T. makuya 10. Elytral tubercles large, often confluent ( Fig. 3F ); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat ( Fig. 3F ) ......................................... T. waclawae - Tubercles medium sized ( Fig. 2E, F ), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex .................................................................................... T. vialis