Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil Author Amorim, Gabriela Dos Santos Programa de Pós-graduação em Biologia Vegetal, Universidade Federal de Pernambuco, Av. Prof. Moraes Rego, 1235 - Cidade Universitária, 50670 - 901, Recife - PE, Brazil. Author Barbosa, Maria Regina De Vasconcellos Departamento de Sistemática e Ecologia, Universidade Federal da Paraíba, Cidade Universitária, Caixa Postal 5065, 58051 - 970, João Pessoa- PB, Brazil. Author Jr, Eduardo Bezerra De Almeida Departamento de Biologia, Universidade Federal do Maranhão, Cidade Universitária Dom Delgado, Av. dos Portugueses, 1966 - Vila Bacanga, 65080 - 805, São Luís - MA. Author Thomas, William Wayt 0000-0002-4996-536X Institute of Systematic Botany, New York Botanical Garden, 2900 Southern Boulevard, Bronx, 10458 - 5126, NY, United States. wthomas @ nybg. org; https: // orcid. org / 0000 - 0002 - 4996 - 536 X wthomas@nybg.org text Phytotaxa 2022 2022-10-11 568 1 27 60 http://dx.doi.org/10.11646/phytotaxa.568.1.3 journal article 163382 10.11646/phytotaxa.568.1.3 87ddb879-1616-4e3b-8817-434b61fb0c60 1179-3163 7184245 11. Eugenia patens Poiret (in Lamarck & Poiret 1813: 124 ). Type :— FRENCH GUIANA . Cayenne , Leblond s.n. (P [01902639] image). ( Fig. 2I , 3 E-H) Trees or shrubs 4 m . Twigs sparsely pubescent, the indumentum reddish-brown. Leaves with petioles 6–9 mm long, pubescent, channeled; blades elliptic to broadly so, or oblong, 10.5–16.2 × 3.8–5.3 cm , glabrous or sparsely pubescent, the indumentum whitish, chartaceous, discolorous; glands conspicuous; base rounded, attenuate or cuneate; apex acute or acuminate; midvein adaxially sulcate; lateral veins visible adaxially, not arched, 19 to 22 pairs; intra-marginal vein ca. 1 mm from the margin, margin flat. Inflorescence axillary, raceme simple, the axis 9.8–12.6 cm long, pubescent; bracts not seen, caducous before anthesis; pedicels 5–15 mm long, pubescent; bracteoles deltate, 8 × 5 mm , pubescent adaxially and glabrous abaxially, caducous after anthesis; flower buds globose ca. 3 mm , not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 2–3 × 3–4 mm , pubescent, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 1–1.3 × 1.3–1.5 cm , pubescent, surface muricate. Note:— Eugenia patens may be recognized by its discolorous, elliptic to broadly elliptic, or oblong leaves, with an adaxially sulcate midvein; and the globose fruit that are pubescent and have a ridged surface. This species is morphologically similar to E . polystachya from which it is distinguished by its fruit with muricate surface (vs. smooth) and glabrous or sparsely pubescent leaves (vs. densely pubescent). Distribution and habitat:— Known from Bolivia , Colombia , Ecuador , French Guiana , Guyana , Peru , Suriname and northeastern and northwestern Brazil ( Acre , Amazonas, Amapá , Maranhão , Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020 ). In Amazonian Maranhão , E. patens was found growing in “terra firme” forests and secondary forests. Phenology:— Flowering and fruiting in October. Specimens examined:— BRAZIL . Maranhão : Bequimão , Pontal , 2°58’54” S , 45°19’09” W , 12 January 2020 (fr.), G.S. Amorim 601 ( JPB !) ; Monção : Ka’apor Indian Reserve , 3°09’58” S , 45°56’17” W , 07 October 1986 (fl.), W. Balée 2684 ( NY !) . Mineirinho , Rio Pindaré , 26 October 1979 (fl., fr), J. Jangoux 937 ( MG !) .