New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region Author Feijoo, Alexander Author Zuluaga, Luis Fernando Author Molina, Ligia Janneth text Zootaxa 2018 2018-10-05 4496 1 448 458 journal article 29251 10.11646/zootaxa.4496.1.34 273e6e1d-05d8-44a5-988d-e7433f986ed8 1175-5326 1446963 ACF076A3-707F-43F5-A121-7CE178A3BB59 Genus Glossodrilus Cognetti, 1905 Andioscolex Michaelsen, 1927 Glossodrilus Righi & Ayres, 1975 . Righi 1975, 1982; Zicsi 1995 . Type species. Glossodrilus parvus Cognetti, 1905 . Diagnosis. Number of species: 55. Distribution: Ecuador (25), Colombia (15), Brazil (5), Costa Rica (4), Panama (4), Venezuela (2), Bolivia (1), Peru (1), Argentina (1). In some cases, the species are distributed in several countries, for example G. nemoralis ( Ecuador , Costa Rica , Panama ), G. parecis ( Argentina , Brazil ), G. crucifer ( Ecuador , Peru ). This list does not include undescribed species listed in Brown & Fragoso (2007) . Genus diagnosis, modified, following Righi (1995) : Eight setae per segment. One pair of intra-clitellar male pores. One pair of calciferous glands in segments 11 and 12 of intertwined-tubular structure, without glandular and membranous division. Three or two pairs of lateral hearts in segments 7–9 and two pairs of intestinal hearts in segments 10, 11. One pair of testes and male funnels present in segment 11. Seminal vesicles generally long. Copulatory chambers absent. Spermathecae mostly present, usually without seminal chambers or diverticula.