Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations Author Williams, Kevin A. Author Cambra, Roberto A. Author Bartholomay, Pedro R. Author Luz, David R. Author Quintero, Diomedes Author Pitts, James P. text Zootaxa 2019 2019-03-05 4564 1 101 136 journal article 28429 10.11646/zootaxa.4564.1.4 6a6ca226-8343-4208-8cdb-f0bd627c3817 1175-5326 2588770 D0A9801B-8049-4211-A4A7-D7792B9D6936 Lomachaeta crocopinna Pitts & Manley, 2004 ( Fig. 52 ) Lomachaeta crocopinna Pitts & Manley, 2004 : 10 . Holotype , ♂ , USA , Texas , Val Verde Co. (UMSP). Pitts & Manley (2004) : Host. Diagnosis. MALE. The following combination of characters is diagnostic for L. crocopinna : the body is entirely black, except the metasoma is almost entirely orange; the mandible lacks a ventral tooth basally; the forewing has its veins encompassing the basal 0.65 × of the wing; the head, mesoscutum, and T2 disc have separated punctures; the T1 shape is sub-sessile; the T2 fringe is composed of slightly thickened setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 3–5 mm . FEMALE. Unknown. Material examined. MEXICO , Sonora : Alamos , Rancho Acosta , malaise in dry wash, tropical deciduous forest, 395 m , 27°01.57’N 106°55.37’W , 26.V–6.VI.2007 , M. E. Irwin ( 1♂ , EMUS ) ; 3 km NE Alamos , La Huerta Orchard , 27°01’N 108°54’W , 19–24.[ Month unknown].2008, M. E. Irwin & O. Figuero ( 2 ♂ , EMUS ) ; USA , Arizona : Yavapal County , 11 mi W. Jerome , Reared, F. D . Parker ( 2♂ , EMUS ) ; Yavapal County , Clarksdale , Reared, F. D . Parker ( 1♂ , EMUS ) ; Coconino County , 7 mi S. Sedona , Reared, F. D . Parker ( 3♂ , EMUS ) ; Gila County , 2 mi S. Pine , F. D. Parker ( 1♂ , EMUS ) ; Pima County , 550m , Silver Reef Wash , 4 km E. Vaiva Vo ( Cockelburr ), Tat Momoli Mts. , malaise in wash, 32°39.06’N 111°54.47’W , 07–14.V.2006 , M. E. Irwin ( 5♂ , EMUS ) ; Maricopa County , Gila River , 10 km S. Arlington , malaise on sand flat, 200m , 33°13.30’N 112°45.53’W , 07–13.VI.2010 , M. E. Irwin ( 1♂ , EMUS ) ; 4 km E. Vaiva Vo , Silver Reef Wash , 32°39.06’N 111°54.47’W , 550m , 1–7.V.2006 , M. E. Irwin , malaise trap ( 1♂ , EMUS ) ; Kitt Peak rd. , Coyote Mts. , 5 km S. jct. Hwy. 36 and 286, malaise in wash, 1300 m , 31°59.32’N 111°33.79’W , 2–12.V.2006 , M. E. Irwin ( 1♂ , EMUS ) ; New Mexico : Socorro , NWR [Natural Wildlife Resort] LTER [Long Term Ecological Reserve] site 1, 26.V.1992 ( 1♂ , EMUS ) ; Texas , Culberson County , Choza Springs , Guadalupe Mountains , 1610 m , 8.IV.1996 ( 1♂ , EMUS , Fig. 52 ) ; Utah : Garfield County , Alvey Wash , 7 km S. Escalante , pan trap in dry wash, 1990 m , 37°42.5’N 111°37.8’W , 24– 25.V.2002 , F. D. Parker & M. E. Irwin ( 2♂ , EMUS ) ; Garfield County , Alvey Wash , 7 km S. Escalante , malaise in dry wash, 1990m , 37°42.5’N 111°37.8’W , 24–25.V.2002 , M. E. Irwin & F. D. Parker ( 1♂ , EMUS ) . Distribution. Known from the Arizona Mountains forests, Colorado Plateau, Great Basin, and Chihuahuan Desert in Arizona , Nevada , New Mexico , and Texas ( USA ) and Sonora ( Mexico ). Remarks. This is the apparent sister species to L. ptilohyalus ; the two species share similar coloration and genitalic morphology. The distribution of L. crocopinna spans multiple ecoregions in the western USA and parallels that of Dilophotopsis concolor (Cresson) . Wilson and Pitts (2008) described the distribution of D. concolor and its sister species, D. paron (Cameron) . Dilophotopsis paron occurs in the Mojave and Sonoran deserts, as does L. ptilohyalus . The parallel distributions of these sister-species pairs should be studied more closely, as they could shed light on historic events that impacted the biogeography of numerous velvet ants.