<document id="44C50E0336906BE1160BB7CC0CB5117C" ID-DOI="10.33910/2686-9519-2022-14-4-570-593" ID-ISSN="2686-9519" ID-Zenodo-Dep="12815734" ID-ZooBank="B3A08C66-2818-4A1D-8834-6751ABFC30A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721869831261" checkinUser="felipe" docAuthor="Sundukov, Yurii N. &amp; Novomodnyi, Evgeniy V." docDate="2022" docId="039F87C8FFD13A6900F6FC69FD03FAD5" docLanguage="en" docName="AmurianZoolJ.XIV.4-570-593.pdf" docOrigin="Amurian Zoological Journal XIV (4)" docSource="http://dx.doi.org/10.33910/2686-9519-2022-14-4-570-593" docStyle="DocumentStyle:1BE7E8521B83181C3ED08D600C3F2F53.3:AmurianZoolJ.2019-.journal_article.english" docStyleId="1BE7E8521B83181C3ED08D600C3F2F53" docStyleName="AmurianZoolJ.2019-.journal_article.english" docStyleVersion="3" docTitle="Elaphrus (Elaphrus) comatus Goulet 1983" docType="treatment" docVersion="2" lastPageNumber="575" masterDocId="FFA6FFB0FFD43A6C0061FE2FFFF7FFC2" masterDocTitle="Ground beetles (Coleoptera, Carabidae) of Khabarovsky Region in the collection of Grodekov Khabarovsk Regional Museum" masterLastPageNumber="593" masterPageNumber="570" pageNumber="575" updateTime="1721876124617" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="429692319E64BDAF0A46540EF70EAA5C">Ground beetles (Coleoptera, Carabidae) of Khabarovsky Region in the collection of Grodekov Khabarovsk Regional Museum</mods:title>
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<mods:namePart id="2B1F9AA9F60DE3965BA4860DF87927B4">Sundukov, Yurii N.</mods:namePart>
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<mods:namePart id="99CBC49A04D74B7C18EBDE5A92C55EC0">Novomodnyi, Evgeniy V.</mods:namePart>
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<treatment id="039F87C8FFD13A6900F6FC69FD03FAD5" LSID="urn:lsid:plazi:treatment:039F87C8FFD13A6900F6FC69FD03FAD5" httpUri="http://treatment.plazi.org/id/039F87C8FFD13A6900F6FC69FD03FAD5" lastPageNumber="575" pageId="5" pageNumber="575">
<subSubSection id="C32C6555FFD13A6900F6FC69FD05FDA6" box="[151,754,582,612]" pageId="5" pageNumber="575" type="nomenclature">
<paragraph id="8B8936DEFFD13A6900F6FC69FD05FDA6" blockId="5.[151,757,582,1303]" box="[151,754,582,612]" pageId="5" pageNumber="575">
<heading id="D0C181B2FFD13A6900F6FC69FD05FDA6" box="[151,754,582,612]" centered="true" fontSize="12" level="2" pageId="5" pageNumber="575" reason="4">
<taxonomicName id="4C364D5DFFD13A6900F6FC69FD05FDA6" ID-CoL="3946Q" authority="Goulet, 1983" authorityName="Goulet" authorityYear="1983" box="[151,754,582,612]" class="Insecta" family="Carabidae" genus="Elaphrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="comatus" subGenus="Elaphrus">
<emphasis id="B942EACCFFD13A6900F6FC69FD05FDA6" bold="true" box="[151,754,582,612]" pageId="5" pageNumber="575">
<emphasis id="B942EACCFFD13A6900F6FC69FEEFFDA1" bold="true" box="[151,280,582,611]" italics="true" pageId="5" pageNumber="575">Elaphrus</emphasis>
(
<emphasis id="B942EACCFFD13A69014BFC69FE5CFDA1" bold="true" box="[298,427,582,611]" italics="true" pageId="5" pageNumber="575">Elaphrus</emphasis>
) 
<emphasis id="B942EACCFFD13A6901DFFC68FDC2FDA1" bold="true" box="[446,565,583,611]" italics="true" pageId="5" pageNumber="575">comatus</emphasis>
Goulet, 1983
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C6555FFD13A6900F6FC42FD21FC21" pageId="5" pageNumber="575" type="materials_examined">
<paragraph id="8B8936DEFFD13A6900F6FC42FD21FC21" blockId="5.[151,757,582,1303]" pageId="5" pageNumber="575">
<emphasis id="B942EACCFFD13A6900F6FC42FEE4FD48" bold="true" box="[151,275,621,650]" pageId="5" pageNumber="575">Material.</emphasis>
<collectingRegion id="49F2F83CFFD13A69014AFC42FE31FD48" box="[299,454,621,650]" country="Russia" name="Khabarovsk" pageId="5" pageNumber="575">Khabarovsk</collectingRegion>
, bank of the 
<materialsCitation id="3B5E3C83FFD13A6902C6FC42FF48FD18" collectingDate="1997-06-14" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Amur">
<collectingRegion id="49F2F83CFFD13A6902C6FC42FD02FD48" box="[679,757,621,650]" country="Russia" name="Amur" pageId="5" pageNumber="575">Amur</collectingRegion>
<location id="8EE96005FFD13A6900F6FCBBFEF1FD73" LSID="urn:lsid:plazi:treatment:039F87C8FFD13A6900F6FC69FD03FAD5:8EE96005FFD13A6900F6FCBBFEF1FD73" box="[151,262,660,689]" name="Channel" pageId="5" pageNumber="575" stateProvince="Amur">Channel</location>
, 
<date id="FF88101EFFD13A690174FCBBFE56FD73" box="[277,417,660,689]" pageId="5" pageNumber="575" value="1997-06-14">
<collectingDate id="EFCCE9F6FFD13A690174FCBBFE56FD73" box="[277,417,660,689]" pageId="5" pageNumber="575" value="1997-06-14">14.06.1997</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A6901CEFCBBFD8AFD73" box="[431,637,660,689]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A6902A4FCBBFD18FD71" box="[709,751,660,691]" count="4" pageId="5" pageNumber="575" type="male">4♂</specimenCount>
, 
<specimenCount id="9D30FD57FFD13A6900F6FC95FF48FD18" box="[151,191,698,730]" count="3" pageId="5" pageNumber="575" type="female">3♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6900B1FC95FF48FCC2" collectingDate="1997-06-30" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Amur">
ibid, 
<date id="FF88101EFFD13A690172FC95FE68FD15" box="[275,415,698,727]" pageId="5" pageNumber="575" value="1997-06-30">
<collectingDate id="EFCCE9F6FFD13A690172FC95FE68FD15" box="[275,415,698,727]" pageId="5" pageNumber="575" value="1997-06-30">30.06.1997</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A6901CFFC94FD8BFD15" box="[430,636,698,727]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A6902A4FC95FD07FD18" box="[709,752,698,730]" count="2" pageId="5" pageNumber="575" type="male">2♂</specimenCount>
, 
<specimenCount id="9D30FD57FFD13A6900F6FCCEFF48FCC2" box="[151,191,737,768]" count="3" pageId="5" pageNumber="575" type="female">3♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6900B1FCCFFF48FCE4" collectingDate="1997-07" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="1" specimenCount-female="1" stateProvince="Amur">
ibid, early 
<date id="FF88101EFFD13A69013DFCCFFE20FD3C" box="[348,471,736,766]" pageId="5" pageNumber="575" value="1997-07">
<collectingDate id="EFCCE9F6FFD13A69013DFCCFFE20FD3C" box="[348,471,736,766]" pageId="5" pageNumber="575" value="1997-07">July 1997</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A690186FCCEFD41FD3F" box="[487,694,736,765]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A6900F6FD28FF48FCE4" box="[151,191,775,806]" count="1" pageId="5" pageNumber="575" type="female">1♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6900B2FD28FD1AFCE4" box="[211,749,775,806]" collectingDate="1998-06-02" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="2" specimenCount-male="2" stateProvince="Amur">
ibid, 
<date id="FF88101EFFD13A690179FD28FE63FCE6" box="[280,404,775,804]" pageId="5" pageNumber="575" value="1998-06-02">
<collectingDate id="EFCCE9F6FFD13A690179FD28FE63FCE6" box="[280,404,775,804]" pageId="5" pageNumber="575" value="1998-06-02">2.06.1998</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A6901C6FD28FD8FFCE6" box="[423,632,775,804]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A6902A7FD28FD1AFCE4" box="[710,749,775,806]" count="2" pageId="5" pageNumber="575" type="male">2♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6900F6FD02FD5EFC8F" box="[151,681,813,845]" collectingDate="1999-07-03" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="1" specimenCount-male="1" stateProvince="Amur">
ibid, 
<date id="FF88101EFFD13A6900BAFD02FEA0FC88" box="[219,343,813,842]" pageId="5" pageNumber="575" value="1999-07-03">
<collectingDate id="EFCCE9F6FFD13A6900BAFD02FEA0FC88" box="[219,343,813,842]" pageId="5" pageNumber="575" value="1999-07-03">3.07.1999</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A690109FD01FDC0FC88" box="[360,567,813,842]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A6902E3FD02FD5EFC8F" box="[642,681,813,845]" count="1" pageId="5" pageNumber="575" type="male">1♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6902DDFD02FD53FCB1" collectingDate="1999-07-03" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Amur">
ibid, 05.2000, 
<collectorName id="26C35308FFD13A690172FD7BFE10FCB3" box="[275,487,852,881]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A69025CFD7BFD90FCB1" box="[573,615,852,883]" count="1" pageId="5" pageNumber="575" type="male">1♂</specimenCount>
, 
<specimenCount id="9D30FD57FFD13A69021CFD7BFD53FCB1" box="[637,676,852,883]" count="1" pageId="5" pageNumber="575" type="female">1♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A6902DDFD7BFDABFC58" collectingDate="2000-06-30" collectorName="E. Novomodnyi" location="Channel" pageId="5" pageNumber="575" specimenCount="1" specimenCount-male="1" stateProvince="Amur">
ibid, 
<date id="FF88101EFFD13A6900F6FD55FED4FC55" box="[151,291,890,919]" pageId="5" pageNumber="575" value="2000-06-30">
<collectingDate id="EFCCE9F6FFD13A6900F6FD55FED4FC55" box="[151,291,890,919]" pageId="5" pageNumber="575" value="2000-06-30">30.06.2000</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A69014CFD54FE01FC55" box="[301,502,890,919]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A690254FD55FDABFC58" box="[565,604,890,922]" count="1" pageId="5" pageNumber="575" type="male">1♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B5E3C83FFD13A690208FD55FF26FC7F" location="Malyshevo" pageId="5" pageNumber="575" specimenCount="1" stateProvince="Amur">
<location id="8EE96005FFD13A690208FD55FD02FC55" LSID="urn:lsid:plazi:treatment:039F87C8FFD13A6900F6FC69FD03FAD5:8EE96005FFD13A690208FD55FD02FC55" box="[617,757,890,919]" name="Malyshevo" pageId="5" pageNumber="575" stateProvince="Amur">Malyshevo</location>
near
</materialsCitation>
<materialsCitation id="3B5E3C83FFD13A6900BCFD8FFD38FC24" collectingDate="1999-05-19" collectorName="E. Novomodnyi" location="Lake Petropavlovskoe" pageId="5" pageNumber="575" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Khabarovsk">
<collectingRegion id="49F2F83CFFD13A6900BCFD8FFE8FFC7F" box="[221,376,928,957]" country="Russia" name="Khabarovsk" pageId="5" pageNumber="575">Khabarovsk</collectingRegion>
, 
<location id="8EE96005FFD13A6901E8FD8FFD51FC7F" LSID="urn:lsid:plazi:treatment:039F87C8FFD13A6900F6FC69FD03FAD5:8EE96005FFD13A6901E8FD8FFD51FC7F" box="[393,678,928,957]" name="Lake Petropavlovskoe" pageId="5" pageNumber="575" stateProvince="Khabarovsk">Lake Petropavlovskoe</location>
, sand spit, 
<date id="FF88101EFFD13A6900B5FDE8FE97FC26" box="[212,352,967,996]" pageId="5" pageNumber="575" value="1999-05-19">
<collectingDate id="EFCCE9F6FFD13A6900B5FDE8FE97FC26" box="[212,352,967,996]" pageId="5" pageNumber="575" value="1999-05-19">19.05.1999</collectingDate>
</date>
, 
<collectorName id="26C35308FFD13A69010BFDE8FDC3FC26" box="[362,564,967,996]" pageId="5" pageNumber="575">E. Novomodnyi</collectorName>
leg., 
<specimenCount id="9D30FD57FFD13A690212FDE8FD6AFC24" box="[627,669,967,998]" count="1" pageId="5" pageNumber="575" type="male">1♂</specimenCount>
, 
<specimenCount id="9D30FD57FFD13A6902C9FDE8FD38FC24" box="[680,719,967,998]" count="1" pageId="5" pageNumber="575" type="female">1♀</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6555FFD13A6900F6FDC2FD03FAD5" pageId="5" pageNumber="575" type="distribution">
<paragraph id="8B8936DEFFD13A6900F6FDC2FD56FB33" blockId="5.[151,757,582,1303]" pageId="5" pageNumber="575">
<emphasis id="B942EACCFFD13A6900F6FDC2FEBAFBC8" bold="true" box="[151,333,1005,1034]" pageId="5" pageNumber="575">Distribution.</emphasis>
A nemoral East Asian species: 
<collectingCountry id="F321764EFFD13A6900B8FA3BFEDBFBF3" box="[217,300,1044,1073]" name="Russia" pageId="5" pageNumber="575">Russia</collectingCountry>
(the south of Khabarovsky Region (
<collectingRegion id="49F2F83CFFD13A6900C1FA15FEC9FB95" box="[160,318,1082,1111]" country="Russia" name="Khabarovsk" pageId="5" pageNumber="575">Khabarovsk</collectingRegion>
) and 
<collectingRegion id="49F2F83CFFD13A6901ECFA15FD77FB95" box="[397,640,1082,1111]" country="Russia" name="Primorskiy" pageId="5" pageNumber="575">Primorsky Region</collectingRegion>
), Northeast 
<collectingCountry id="F321764EFFD13A6900B3FA4FFED6FBBF" box="[210,289,1120,1149]" name="China" pageId="5" pageNumber="575">China</collectingCountry>
(
<collectingRegion id="49F2F83CFFD13A690150FA4FFE29FBBF" box="[305,478,1120,1149]" country="China" name="Heilongjiang" pageId="5" pageNumber="575">Heilongjiang</collectingRegion>
); also listed from 
<collectingCountry id="F321764EFFD13A6902A2FA4EFF37FB61" name="South Korea" pageId="5" pageNumber="575">Korea</collectingCountry>
(National Institute of Biological Resources 2019), Hokkaido (
<bibRefCitation id="EFA74B2FFFD13A6901E9FA82FD5EFB08" author="Kimoto, Sh. &amp; Yasuda, N." box="[392,681,1197,1226]" pageId="5" pageNumber="575" refId="ref17375" refString="Kimoto, Sh., Yasuda, N. (1995) Carabidae in Hokkaido. Tokyo: Tokai University Press, 317 p. (In Japanese)" type="book" year="1995">Kimoto, Yasuda 1995</bibRefCitation>
), and South Sakhalin (
<bibRefCitation id="EFA74B2FFFD13A69010AFAFBFD63FB33" author="Yoshimatsu, S. - I. &amp; Ito, N. &amp; Nakatani, Y. &amp; Yoshitake, H." box="[363,660,1236,1265]" pageId="5" pageNumber="575" pagination="15 - 191" refId="ref20823" refString="Yoshimatsu, S. - I., Ito, N., Nakatani, Y., Yoshitake, H. (2018) A list of ground beetles (Insecta: Coleoptera: Caraboidea) in Dr. Kazuo Tanaka collection preserved in the Insect Museum of Institute for Agro-Environmental Sciences, NARO. Bulletin of the NARO, Agro-Environmental Sciences, vol. 39, pp. 15 - 191. (In English)" type="journal article" year="2018">Yoshimatsu et al. 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8B8936DEFFD13A6900F6FAD5FD03FAD5" blockId="5.[151,757,582,1303]" box="[151,756,1274,1303]" pageId="5" pageNumber="575">
<emphasis id="B942EACCFFD13A6900F6FAD5FF16FAD4" bold="true" box="[151,225,1274,1302]" pageId="5" pageNumber="575">Note.</emphasis>
First reported for the Khabarovsky Region.
</paragraph>
</subSubSection>
</treatment>
</document>