Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes
Author
Gusarov, Vladimir I.
text
Zootaxa
2003
2003-11-17
353
1
1
134
https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
journal article
5458
10.11646/zootaxa.353.1.1
740af13a-ed5c-40d0-b0c1-8d3fd2b92f56
11755334
5019783
Atheta
(
s. str.
)
graminicola
(
Gravenhorst, 1806
)
(
Fig.
155
in
Strand & Vik 1964
)
Aleochara graminicola
Gravenhorst, 1806: 176
.
Homalota granulata
Mannerheim, 1846: 508
(misspelled as
Stonalota granulata
).
Liogluta
(
Megista
)
graminicola
:
Mulsant & Rey, 1874a: 624
(as valid species).
Atheta
(
Megista
)
graminicola
:
Bernhauer, 1907: 394
(as valid species).
Atheta graminicola
:
Fenyes, 1907: 61
(as valid species).
Atheta granulata
:
Fenyes, 1907: 61
(as synonym of
At. graminicola
).
Atheta
(
Homalotusa
)
lanei
Casey, 1910a: 10
,
syn. nov.
Atheta
(
Megista
)
nomadica
Casey, 1910a: 15
,
syn. nov.
Atheta
(
Megista
)
graminicola
:
Bernhauer & Scheerpeltz, 1926: 659
(as valid species).
Atheta
(
Megista
)
granulata
:
Bernhauer & Scheerpeltz, 1926: 659
(as synonym of
At. graminicola
).
Atheta
(
Megista
)
nomadica
:
Bernhauer & Scheerpeltz, 1926: 659
(as valid species).
Atheta
(
Homalotusa
)
lanei
:
Bernhauer & Scheerpeltz, 1926: 655
(as valid species).
Atheta
(
s. str.
)
graminicola
:
Strand & Vik, 1964: 155
(as valid species).
Atheta
(
s. str.
)
graminicola
:
Benick & Lohse, 1974: 195
(as valid species).
Atheta graminicola
:
Lohse & Smetana, 1985: 286
(as valid species).
Atheta granulata
:
Lohse & Smetana, 1985: 284
(as valid species).
Atheta graminicola
:
Lohse
et al.
, 1990: 190
(as valid species).
Atheta granulata
:
Lohse
et al.
, 1990: 189
(as valid species).
Atheta nomadica
:
Lohse
et al.
, 1990: 189
(as synonym of
At. granulata
).
(Other references are omitted)
Type material.
Lectotype
of
Aleochara graminicola
(here designated):
♂
, “5349”, “
graminicola
Gr.
”, “Zool. Mus.
Berlin
” (
ZMHB
)
.
Paralectotypes
:
2♂♂
,
2♀♀
, without labels;
♀
, “Angl. Rudd”
;
♀
, “Nordeur. Schupp.” (
ZMHB
)
.
Lectotype
of
Homalota granulata
(designated by
Lohse and Smetana (1985))
:
UNITED STATES
:
Alaska
:
♀
,
Unalaska Island
(Cygnaeus) (
MZHF
).
Lectotype
of
Atheta lanei
(here designated):
♀
, “
Lane Co.
, Or.[egon]”, “
lanei
Csy.
”, “TYPE USNM 39462” (red label), “CASEY bequest 1925” (
NMNH
).
Lectotype
of
Atheta nomadica
(here designated):
♀
, “
Nome
,
Alaska
”, “
nomadica
Csy.
”, “TYPE USNM 39478” (red label), “CASEY bequest 1925” (
NMNH
).
Additional material.
CANADA
:
Alberta
:
♂
,
♀
, Banff (Fenyes);
3♀♀
,
2 mi.
S Ponaka, Battle Riv. marsh,
6.viii.1976
(P.M.Hammond);
♂
,
2♀♀
, Waterton N. P., Cameron Lake, 540’,
2–4.viii.1976
(P.M.Hammond) (
BMNH
);
British Columbia
:
♂
, Alta Lake,
1.viii.1975
(J.M. & B.A.Campbell) (
CNCI
);
Labrador:
♀
, L’Anse au Loup,
9.viii.1972
(J.M.Campbell);
2♂♂
, Hebron,
9.vii.1954
(J.F.McAlpine) (
CNCI
);
Manitoba
:
♀
, Churchill,
29.vi.1937
(W.J.Brown);
♀
, ditto but
20.vii.1937
;
♂
, ditto but
18.viii.1937
;
♂
,
2♀♀
, ditto but
16.vi.1937
;
2♂♂
,
6♀♀
, ditto but
4.vi.1952
(J.G.Chillcott);
4♂♂
,
4♀♀
, ditto but
13.vi.1952
;
♂
,
3♀♀
, ditto but
17.vi.1952
;
♂
,
♀
, ditto but
2.viii.1952
;
3♂♂
,
♀
, ditto but
5.viii.1952
;
♂
, ditto but
15.viii.1952
;
♀
, ditto but
27.viii.1948
(W.R.Richards) (
CNCI
);
Newfoundland:
♂
, Long Range Mts.,
50º34’N
56º56’W
, 1400’,
24.vii.1969
(M.J.D.Brendell) (
BMNH
);
Northwest Territories
:
2♀♀
, Ft. Simpson, Harris River,
15.vi.1972
(A.Smetana);
♂
, ditto but
10.vi.1972
;
5♂♂
,
19♀♀
, Trail River,
32 mi.
NW Ft. Simpson,
16.vi.1972
(A.Smetana);
3♂♂
,
4♀♀
, Hwy.
3, 5 mi.
SE Ft. Simpson,
17.vi.1972
(A.Smetana);
3♂♂
,
2♀♀
, ditto but
21.vi.1972
;
♀
, Hwy.
3, 8 mi.
SE Ft. Simpson,
18.vi.1972
(A.Smetana);
♂
, Spence River,
38 mi.
SE Ft. Simpson,
19.vi.1972
(A.Smetana);
3♂♂
,
13♀♀
, Reindeer Sta., Caribou Hills,
30.vi.1972
(A.Smetana);
♀
, Yellowknife,
20.v.1953
(J.G.Chillcott);
♂
, ditto but
29.v.1953
;
2♀♀
, Aklavik,
9.vi.1956
(R.E.Leech);
2♀♀
, ditto but
12.vi.1956
;
♂
, ditto but
14.vi.1956
;
♂
, ditto but
3.vii.1956
;
♀
, ditto but
6.vii.1956
;
♂
, ditto but
13.vi.1956
(E.F.Cashman);
♀
, ditto but
16.vi.1953
(C.D.Bird);
10♀♀
, Inuvik, Shell Lk.,
27.vi.1972
(A.Smetana);
♀
, ditto but
1.vii.1972
(A.Smetana);
6♀♀
, ditto but
25.vi.1972
;
2♂♂
,
♀
, Reindeer Depot, Mackenzie Delta,
8.viii.1948
(W.J.Brown);
♀
, Kidluit Bay, N. Richards Is.,
26.vii.1948
(T.J.R.Vockeroth);
♀
, Muscox L.,
64º45’N
108º10’W
,
2.viii.1953
(J.G.Chillcott);
3♂♂
,
1 specimen
, ditto but
21.vii.1953
(J.G.Chillcott);
♂
, Salmita Mines,
64º05’N
111º15’W
,
20.vi.1953
(J.G.Chillcott);
2♀♀
, Unnamed Lake,
18 mi.
NW Inuvik via East Channel,
26.vi.1972
(A.Smetana) (
CNCI
);
Ontario
:
♂
,
2♀♀
, Butterfly Lk.,
13 mi.
S Sioux Lookout,
17.vi.1973
(Campbell & Parry) (
CNCI
);
Québec
:
7♂♂
,
15♀♀
, Blanc Sablon,
9.viii.1972
(J.M.Campbell);
♂
,
2♀♀
, Koroc Riv., in leaf litter beside stream,
3.viii.1986
(A.V.A. & S. Morgan);
♂
, Whale Riv.,
8.vii.1949
(J.R.Vockeroth) (
CNCI
);
Yukon Territory
:
♀
, Caribou Bar Creek,
20.vi.1972
;
2♂♂
,
5♀♀
, McQuesten Lk.,
15 mi.
SW Keno,
19.vii.1968
(Campbell & Smetana);
2♂♂
,
3♀♀
, Mi. 1034, Alaska Hwy., near Kloo Lake,
5.vii.1968
(Campbell & Smetana);
4♀♀
, Mi. 1120, Alaska Hwy.,
6.vii.1968
(Campbell & Smetana);
♀
, Mi. 1192, Alaska Hwy., near Snag Junction,
6.vii.1968
(Campbell & Smetana);
♀
, Mi. 1120, Alaska Hwy.,
6.vii.1968
(Campbell & Smetana);
♂
,
♀
, Mi. 1705, Alaska Hwy., Slim’s River,
7.vii.1981
(Bright);
♂
,
10 mi.
E Dawson City,
11.vii.1968
(Campbell & Smetana);
3♂♂
,
2♀♀
, Dempster Hwy., Mi. 236, Eagle Riv., 1400’,
12.vii.1978
(Campbell & Smetana);
2♂♂
, Dempster Hwy., Mi. 222, 1900’,
21.vii.1978
(Smetana & Campbell);
3♂♂
,
6♀♀
, ditto but Mi. 147,
22.vii.1978
(Campbell & Smetana);
2♀♀
, ditto but Mi. 139.5,
20.vii.1978
(Campbell & Smetana);
♀
, ditto but Mi. 73, 3300’,
12.vii.1968
(Campbell & Smetana);
2♂♂
,
10♀♀
, ditto but Mi. 71;
♂
,
7♀♀
, ditto but Mi. 65, 3300’,
13.vii.1968
;
4♂♂
,
♀
, ditto but Mi. 60, 3500’,
19.vii.1978
;
4♂♂
,
4♀♀
, ditto but
23.vii.1978
;
♂
,
2♀♀
, ditto but Mi. 55, 3300’,
13.vii.1968
;
5♂♂
,
2♀♀
, ditto but Mi. 53, North Fork Pass, 4200’,
24.vii.1978
;
3♂♂
,
♀
, ditto but Mi. 48.5, North Fork Pass, 4100’,
24.vii.1978
;
♂
,
3♀♀
, ditto but
19.vii.1978
;
5♂♂
,
2♀♀
, ditto but Mi. 45, 3500’,
14.vii.1968
;
♀
, ditto but 3400’,
18–23.vii.1978
(J.M.Campbell);
11♂♂
,
13♀♀
, ditto but Mi. 42, N Klondike Riv., 3300’,
18.vii.1978
(Campbell & Smetana);
♂
, ditto but Mi. 32, 2700’,
15.vii.1968
;
♂
, ditto but Mi. 28,
15.vii.1968
;
♀
, Ogilvie Mts., North Fork Pass, 4100’,
18.vi.1962
(P.J.Skitsko);
♂
,
♀
, Rampart House,
28.v.1951
(J.E.H.Martin);
♂
, ditto but
1.vi.1951
;
4♀♀
, British Mts.,
340 m
,
4 km
NE Wolf Cr.,
69º08’N
140º14’W
,
23.vi.1984
(Campbell);
♂
,
♀
, British Mts.,
320 m
,
3 km
NW Firth Riv.,
69º13’N
140º05’W
, sifting
Salix
litter,
19.vi.1984
(Campbell);
♂
,
♀
, British Mts.,
180 m
, Fish Creek,
69º27’N
140º19’W
, sifting under tall
Salix
shrubs,
3.vii.1984
(J.M.Campbell);
2♂♂
,
3♀♀
, Hansen Lk.,
9 mi.
SW Keno,
18.vii.1968
(Campbell & Smetana);
♂
, La Force Lake,
62º41’N
132º20’W
, 3300’,
28.vi.1960
(J.E.H.Martin);
♀
, Swim Lakes,
62º13’N
133º00’W
, 3200’,
15.vi.1960
(J.E.H.Martin) (
CNCI
);
♀
, Whitehorse,
21.vi.1924
(
MCZ
);
UNITED STATES
:
Alaska
:
16♂♂
,
17♀♀
, Mi. 123, Taylor Hwy.,
9.vii.1968
(Campbell & Smetana);
♀
, Mi. 23, Taylor Hwy.,
8.vii.1968
(Campbell & Smetana);
♀
, Denali State Park, Mi. 165.5, Hwy. 3,
23.vi.1978
(Smetana & Becker);
2♂♂
,
3♀♀
, Mi. 110, Denali Hwy., Seattle Cr.,
15.vii.1978
(Campbell & Smetana);
4♂♂
,
5♀♀
, Mi. 102, Denali Hwy.,
12.vii.1978
(Smetana & Campbell);
2♀♀
, Mi. 94, Denali Hwy., Canyon Creek,
13.vii.1978
(J.M.Campbell & A.Smetana);
♂
,
12 mi.
N Mi. 78, Denali Hwy., Windy Cr.,
13.vii.1978
(Smetana & Campbell);
6♂♂
,
8♀♀
, Mi. 1249, Alaska Hwy., Deadman Lk.,
6–7.vii.1968
(Campbell & Smetana);
2♂♂
,
6♀♀
, Mi. 1252, Alaska Hwy.,
7.vii.1968
(Campbell & Smetana);
2♂♂
,
♀
, Kenai Pen., Kenai,
24.vi.1951
(W.J.Brown);
♂
,
♀
, Kenai Pen.,
3 mi.
SE Kenai,
7.vi.1978
(Smetana & Becker);
♀
, Kenai Pen., Hope,
24.v.1951
(W.J.Brown);
5♂♂
,
7♀♀
, Kenai Pen., Anchor River at Hwy. 1, 450’,
2.vi.1978
(Smetana & Becker);
♂
,
20♀♀
, ditto but
4.vi.1978
;
♀
, ditto but
5.vi.1978
;
♂
,
4♀♀
, Kenai Pen., Anchor River Campground,
12 mi.
N Homer, 450’,
5.vi.1978
(Smetana & Becker);
♀
, ditto but
3.vi.1978
;
4♂♂
,
5♀♀
, Kenai Pen.,
1 mi.
W Anchor Pt.,
1.vi.1978
(Smetana & Becker);
7♂♂
,
3♀♀
, Kenai Pen., Crooked Cr. at Hwy. 1, S Kasilof,
9.vi.1978
(Smetana & Becker);
5♂♂
,
11♀♀
, Kenai Mts.,
15 mi.
N Seward, 400’,
29.v.1978
(Smetana & Becker);
6♂♂
,
9♀♀
, Kenai Mts.,
16 mi.
N Seward, 500’–600’,
26.v.1978
(Smetana & Becker);
4♂♂
,
6♀♀
, Kenai Mts.,
22 mi.
N Seward, 600’–800’,
27.v.1978
(Smetana & Becker);
2♂♂
,
5♀♀
, Seward,
20.v.1951
(W.J.Brown);
2♂♂
, Kenai Pen.,
15 mi.
E Sterling,
14.vi.1978
(Smetana & Campbell);
2♂♂
,
♀
, Kenai Pen.,
2 mi.
NE Soldotna,
10.vi.1978
(Smetana & Becker);
4♂♂
, Kenai Pen., Clam Gulch,
6.vi.1978
(Smetana & Becker);
3♂♂
,
5♀♀
, Kenai Mts., Cooper Lk., 1000’,
15.vi.1978
(Smetana & Becker);
♀
, Kenai Mts.,
8 mi.
W Cooper Landing, 500’,
14.vi.1978
(Smetana & Becker);
♀
, Kenai Mts., Quarts Cr. At Hwy. 1, 500’,
15.vi.1978
(Smetana & Becker);
♂
,
3♀♀
, Kenai Mts., Tern Lk. Campground, 700’,
18.vi.1978
(Smetana & Becker);
♀
,
8 mi.
NW Haines,
3.vii.1968
(Campbell & Smetana) (
CNCI
);
3 specimens
, Skagway,
18– 20.vi.1924
(H.C.Fall) (
CUIC
);
4♂♂
,
♀
, ditto but
18.vi.1924
(
MCZ
);
3♂♂
,
8♀♀
, Hess Cr., Mi. 24, Wales Hwy.,
1.vii.1978
(J.M.Campbell & A.Smetana);
3♂♂
,
♀
, ditto but 600’,
65º40’N
149º10’W
,
10.vii.1978
(J.M.Campbell & A.Smetana);
9♂♂
,
5♀♀
, Unalakleet,
4.vii.1961
(R.Madge);
2♀♀
, ditto but
23.vi.1961
;
♀
, ditto but
17.vi.1961
;
♀
, ditto but
11.viii.1961
;
♀
, King Salmon, Naknek River,
6.vii.1952
(W.R.Mason);
1 specimen
, Nome,
10.vii.1951
(D.P.Whillians);
♂
, Mi. 220, George Parks Hwy.,
14.vii.1978
(A.Smetana) (
CNCI
);
2♀♀
, Fairbanks,
7.vii.1924
(
MCZ
);
2♀♀
, Prudhoe Bay Rd., Bonanze Cr.,
66º40’N
150º40’W
, 900’,
2.vii.1978
(A.Smetana & J.M.Campbell);
♀
, Prudhoe Bay Rd.,
4 mi.
SW Chandalar Cmp.,
68º04’N
149º40’W
, 4000’,
7.vii.1978
(Campbell & Smetana);
2♂♂
,
4♀♀
, Prudhoe Bay Rd., Coldfoot nr. Clara Cr.,
67º16’N
150º10’W
, 1100’,
3.vii.1978
(A.Smetana & J.M.Campbell);
3♀♀
, Prudhoe Bay Rd.,
8 mi.
N S. Fork Koyukuk Riv.,
67º13’N
150º07’W
, 1000’,
8.vii.1978
(A.Smetana & J.M.Campbell);
4♂♂
,
6♀♀
, Prudhoe Bay Rd., 0.5 mi. N Yukon Riv.,
65º52’N
149º45’W
, 400’,
9.vii.1978
(A.Smetana & J.M.Campbell) (
CNCI
);
2 specimens
, Glenn Hwy. (Tok cutoff), NE of Gakona, 62º19.89–.88'N 145º12.60–.27'W,
550 m
, a bog, lake shore,
Carex
,
Comarum
,
16.vii.1998
(V.I.Gusarov);
4 specimens
, Kenai Peninsula,
3 km
SE Anchor Point, Sterling Hwy., Anchor River valley,
59º45.40'N
151º47.61'W
, 0 m, forest litter,
Alnus
,
22.vii.1998
(V.I.Gusarov) (
SPSU
);
♀
, Dutch Harbor Unalaska I.,
9.vii.1907
(E.C.Van Dyke) (
MCZ
);
RUSSIA
:
Chukotka
Pen.:
7 specimens
, NW part of the Krest Bay, along river,
Salix
grove with dense herbs at the margins,
28.vii.1988
(Yu.M.Marusik) (
SPSU
);
Moscow
Reg.:
♂
, Ramenskoye distr., Kraskovo, river bank,
30.vi.1989
(V.I.Gusarov) (
SPSU
);
Murmansk
Reg.:
10 specimens
,
150 km
NE Murmansk, env. of Dalniye Zelentsy, Barentsevo Sea, Yarnyshnaya, seashore, under beach wrack,
4.viii.1997
(A.Przhiboro);
5 specimens
, Khibiny Mts., Bolshoy Vud'yavr Lake,
31.vii.1984
(V.I.Gusarov) (
SPSU
).
Diagnosis.
See
Benick & Lohse (1974)
. Aedeagus and spermatheca:
Fig.
155
in
Strand & Vik (1964)
.
Discussion.
The
lectotype
of
At. nomadica
represents a small specimen of
At. graminicola
, and falls within the range of intraspecific variation of
At. graminicola
in punctation, proportions of antennal segments and the shape of the spermatheca. The
lectotype
of
At. lanei
is similar to examined specimens of
At. graminicola
in external characters and in the shape of the spermatheca.
Lohse and Smetana (1985)
argued that
At. graminicola
and
At. granulata
are closely related but different species, citing the difference in the shape of the aedeagus and spermatheca and in punctation of the pronotum and elytra. The proximal portion of the spermatheca in
At. graminicola
is variable and the difference between
Figs. 5 and
7
in
Lohse and Smetana (1985)
is within the range of variability of this species. I was unable to confirm the difference between Palaearctic specimens of
At. graminicola
and Nearctic specimens. I consider
At. granulata
to be synonymous with
At. graminicola
.
Distribution.
Atheta graminicola
is a Holarctic species. In North America it is known from
Canada
(
British Columbia
,
Yukon Territory
,
Northwest Territories
,
Alberta
,
Manitoba
,
Ontario
,
Québec
, Labrador, Newfoundland) and the
United States
(
Alaska
and
Oregon
).