3473
Author
Published, First
text
Zootaxa
2012
2012-09-07
3473
1
60
journal article
11755334
3D1227F1-4953-4EEA-8600-FE1F117D1B08
Coluber melanzostus
Gravenhorst, 1807
Coluber melanzostus
Gravenhorst, 1807: 402
.—Type locality, restricted by virtue of
neotype
designation: “Batavia,
Java
”, now
Jakarta
,
Java
Island
,
Indonesia
.—Onomatophore.
Neotype
, by present designation:
MNHN 0060
.
Status. A
valid taxon for populations of
Indonesia
(
Java
,
Bali
and
Sumatra
)
.
Comments.
Boie (1826
: col. 206) provided a detailed description of this taxon. It was synonymized with
Tropidonotus quincunciatus
by
Schlegel (1837: 308)
but was recognized at a subspecific rank by several authors (see
Smith 1943
). It was raised to full species status by
Das (1996)
, a position confirmed by Vogel & David (2007).
Gravenhorst (1807: 402–403)
described three varieties, identified by letters “a” to “c”. Obviously, he had at hand at least
three syntypes
. No type locality was given in the original description. It was subsequently stated to be “
Java
” by
Boie (1826
: col. 206). The three or more
syntypes
were deposited in the Museum of Bratislava. They are at present untraced and were obviously destroyed. According to W. Böhme (pers. comm.,
January 2009
), Gravenhorst was working in Bratislava; the collection his book was based upon was later given to this museum (
Adler 1989
). The largest part of the old collection of Bratislava was destroyed during the Second World War.
We consider
Xenochrophis melanzostus
a variable species, containing either striped or blotched individuals. In order to associate the specific nomen
melanzostus
with the striped phase, we design a
neotype
for
Coluber melanzostus
Gravenhorst,
1807
in order to fix the nomenclatural status of this taxon. In agreement with the various dispositions of Article 75.3 of the
Code
and based on (1) the need to fix the status of
Coluber melanzostus
Gravenhorst, 1807
and associate this combination with Indonesian specimens showing a striped dorsal pattern, (2) the destruction of the
syntypes
, which are definitely not present in museums according to the curator of the collection of Bratislava, (M. Rybecký in litt.) (3) the morphological similarity of the selected
neotype
with one of the phase described by Gravenhorst, (4) the origin of the
neotype
from the same locality as the
syntypes
, we designate a
neotype
of
Coluber melanzostus
Gravenhorst.
Neotype
(
Figs. 2–4
):
MNHN 0060
, an adult female from “Batavia”, now
Jakarta
,
Java
Island,
Indonesia
. Collected by
Mr. Méder.
Morphology and measurements
. The body is elongate but robust; head elongate, moderately distinct from the thick neck; snout long, narrowing anteriorly, about 1.8 times eye diameter, without distinct canthus rostralis; nares small, crescentic, piercing upper half of nasal, directed obliquely upwards; eye large, diameter about 1.5 times eyelip distance, round pupil; tail average for the genus, thick, tapering. SVL
605 mm
, Tal
200 mm
, TL
805 mm
; ratio TaL / TL 0.248; HL
33.6 mm
.
Maxillary teeth.
On left, 25 teeth, posterior ones only slightly larger than anterior ones, and row continuous.
FIGURE 2.
MNHN 0060,
Xenochrophis melanzostus
, female, from Jakarta, Java, Indonesia. Neotype of
Coluber melanzostus
Gravenhorst, 1807
. TL 805 mm. Dorsal view. Photo P. David.
FIGURE 3
. MNHN 0060,
Xenochrophis melanzostus
, female, from Jakarta, Java, Indonesia. Neotype of
Coluber melanzostus
Gravenhorst, 1807
. TL 805 mm. Ventral view. Photo P. David.
Scalation
. DSR 19–19–17 (first reduction at left at
VEN
77), distinctly keeled except for the two ventralmost scale rows on each side; 1st DSR much enlarged;
VEN
139 (+ 2 pVEN); SC 70, all paired; anal divided. Rostral wider than high, well visible from above; nasals large, subrectangular, divided below nostril; internasals small, triangular, distinctly narrowing anteriorly, as long as prefrontals; prefrontal very short, 0.4 time as long as frontal; frontal pentagonal, about 1.8 longer than wide; parietals long and wide, 1.2 times as long as frontal; 1 / 1 loreal small, slightly higher than long; 9 / 9 SL, SL
1–2 in
contact with nasal,
2–3 in
contact with loreal, SL 4–5 entering orbit, SL 8 largest; 1 / 1 PreOc, 3 / 3 PostOc, no subocular; temporals 2+3 / 2+3, lower anterior one large and elongate, upper posterior one in broad contact with parietal; 10 / 10 IL, first pair in contact, IL
1–5 in
contact with anterior chin shields; anterior chin shields shorter than posterior shields.
FIGURE 4.
MNHN 0060,
Xenochrophis melanzostus
, female, from Jakarta, Java, Indonesia. Neotype of
Coluber melanzostus
Gravenhorst, 1807
. TL 805 mm. Lateral view of the head. Photo P. David.
Colour and pattern
(in preservative). Dorsal surface of body and tail pale yellowish-grey, with many scales of DSR 5–6 (counted from the ventrals) edged with dark brown, brighter and paler on lower half of DSR 2, with a pair of conspicuous, longitudinal chestnut brown stripes extending from neck to base of tail on each side, as follows: a broad stripe on upper half of DSR 2, DSR 3 and lower half of DSR 4, becoming progressively narrower posteriorly and eventually covering only upper half of DSR 2 and lower half of DSR 3; a narrower stripe on upper half of DSR 6 and all of DSR 7, broad and solid anteriorly but quickly narrowing and coloured in its middle with pale yellowish-grey on much of DSR 7, so as to appear as two narrow, parallel brown lines separated by a pale yellowish-grey area, especially posteriorly; a vertebral stripe, entirely pale yellowish-grey anteriorly, briefly edged with dark chestnut-brown and appearing as two narrow chestnut-brown stripes widely separated by a narrow, irregular pale yellowish-grey stripe; lower posterior margin of scales of DSR 1 clouded with dark grey. Tail as body above but with various chestnut-brown lines reduced spots or dark edges of scales.
Head greyish-olive, darker than body, nearly uniformly coloured except lower part of supralabials, somewhat paler, and of a very faint, dark brown oblique marking on top and posterior edge of SL 6 below each eye; a conspicuous sagittal, elongate cream, brown-edged streak just behind limit of parietal; a conspicuous, vertical chestnut-brown crossbar behind posterior end of upper jaw, connected to upper dorsal stripe and not reaching ventrals, which look like a widely open “V” when see from above.
Chin
and throat uniformly pale yellowish-grey. Venter pale yellowish-grey, tips and anterior margins of ventrals with dark brown margins, producing conspicuous crossbars. Tail as venter with narrow, dark chestnut-brown crossbars.