Systematics of Phasia Latreille (Diptera: Tachinidae) Author Sun, Xuekui Author Marshall, Stephen A. text Zootaxa 2003 2003-08-29 276 1 1 320 https://biotaxa.org/Zootaxa/article/view/zootaxa.276.1.1 journal article 10.11646/zootaxa.276.1.1 1175­5334 5019510 4177F6E7-B276-438E-8E49-EE5ECD84D351 3.2. The Phasia argentifrons species­group The Phasia argentifrons species­group includes all species previously placed in the subgenus Mormonomya Brauer and Bergenstamm ( Crosskey 1973 , 1984 , Cantrell and Crosskey 1989 ), the genus Epaulophasia Townsend ( Townsend 1934 , Guimarães 1971 ) and the genus Campbellia Miller (Miller 1932, Dugdale 1969 , Cantrell and Crosskey 1989 ). Diagnosis : Head compressed anteriorly or spherical; fronto­orbital plate entirely bare laterally, not swollen; dorsal facets of eyes larger than ventral ones; inner margins of eye divergent. Parafacial bare; lower margin of face strongly projecting, almost forming a right angle with face. Sublunular bulla spherical or knob­like; first flagellomere short. Length of oral opening more than 2.5 times width; occipital hairs white. Thorax black; scale­like setae sometimes present in male; scutellum black; subscutellum exceptionally prominent. M meeting R 4+5 at right angle. Abdomen subovate. Male surstylus longer than cercus; phallus short, haired or bare; apex of pregonite broad; postgonite well developed. Key to species of the Phasia argentifrons species­group (The males of Phasia jeanneli , Phasia furcata and Phasia nigromaculata and the females of Phasia distincta and Phasia brachyptera are unknown. Unrevised [Neotropical] species are marked with an asterisk) 1. Female ....................................................................................................................... 2 ­ Male ........................................................................................................................ 10 2. Apex of sternite VII bent or directed ventrally ......................................................... 3 ­ Apex of sternite VII straight or directed dorsally ..................................................... 8 3. Sternite VII broad, strongly bent downward posteriorly .......................................... 4 ­ Sternite VII narrow, only slightly bent downward .................................................... 6 4. Lower margin of face orange yellow; posterior margin of sternite VII almost spherical, with a shallow median notch............................. P. argentifrons Walker, 1849 ♀ ­ Lower margin of face brown, or black; posterior margin of sternite VII not as above .................................................................................................................................... 5 5. Sternite VII branched ................................................ P. furcata Sun , new species ♀ ­ Sternite VII pointed (Fig. I­32.1). ............................ P. lepidofera ( Malloch, 1929 ) ♀ 6. Sternite VII laterally flattened in apical half, narrowest at middle in profile (Fig. I­13.6) ...................................................................................... P. campbelli ( Miller, 1923 ) ♀ ­ Sternite VII not as above ........................................................................................... 7 7. Apex of sternite VII distinctly knobbed in profile (Fig. I­39.7) ................................. ............................................................................................ P. nasuta ( Loew, 1852 ) ♀ ­ Apex of sternite VII straight in profile (Fig. I­27.3)..... P. jeanneli ( Mesnil, 1953 ) ♀ 8. Sternite VII arrow shaped in ventral view (as in Fig. I­42.6) ..................................... .............................................................................. * P. officialis ( Townsend, 1934 ) ♀ ­ Sternite VII broadened near apex, with a straight posterior margin in ventral view (Fig. I­34.6) .................................................... P. nigromaculata Sun , new species ♀ ­ Sternite VII tapered in ventral view .......................................................................... 9 9. Sternite VII slightly flattened at base (Afrotropical) .................................................. ..................................................................... P. nigrofimbriata ( Villeneuve, 1935 ) ♀ ­ Sternite VII not flattened (Australasian and Pacific) .... P. sensua ( Curran, 1927 ) ♀ ­ Sternite VII flattened in apical half (Neotropical) ..................................................... ................................................................................ * P. capitata ( Townsend, 1927 ) ♀ 10. Lower margin of face orange yellow ...................................................................... 11 ­ Lower margin of face brown to black ..................................................................... 12 11. Distiphallus well sclerotized; phallus not haired; syncercus deeply notched posteriorly (Fig. I­6.3­4) ..................................................... P. argentifrons Walker, 1849 ♂ ­ Distiphallus membranous; phallus sparsely haired dorsally; syncercus with a shallow notch posteriorly (Fig. I­13.4­5) .......................... P. campbelli ( Miller, 1923 ) ♂ 12. Parafacial 2.8 times as wide as first flagellomere; both basiphallus and distiphallus well sclerotized (Fig. I­15.6); wing base and hind femur with scale­like setae........... .................................................................................. P . distincta Sun , new species ♂ ­ Parafacial less than 2.0 times as wide as first flagellomere; basiphallus usually membranous; wing base and hind femur with or without scale­like setae ..................... 13 13. Abdomen with yellow spot or area laterally; ejaculatory apodeme fan­like (Fig. I­12.4) ............................................................................ P. brachyptera Sun , new species ♂ ­ Abdomen black; ejaculatory apodeme not fan­like ............................................... 14 14. Phallus haired dorsally ............................................................................................ 15 ­ Phallus not haired .................................................................................................... 16 15. Syncercus slender, with a deep U­shaped notch posteriorly; surstylus strongly arched ........................................................................................ P. sensua ( Curran, 1927 ) ♂ ­ Syncercus broad and short, with a V­shaped notch posteriorly; surstylus slightly arched ................................................................................. P. lepidofera ( Malloch, 1929 ) ♂ 16. Syncercus with a pair of teeth posteriorly, phallapodeme not strongly arched....... 17 ­ Syncercus usually without teeth posteriorly; if teeth present, phallapodeme strongly arched ............................................................................................................... 18 17. Teeth of syncercus situated in the middle of notch; distiphallus partly sclerotized .... .................................................................................* P. capitata ( Townsend, 1927 ) ♂ ­ Teeth of syncercus situated at posterior margin of notch; distiphallus membranous (Fig. I­39.3­4) .................................................................... P. nasuta ( Loew, 1852 ) ♂ 18. Phallapodeme strongly arched; notch of syncercus trapezoid, with two small teeth in middle (Fig. I­41.3­4)................................... P. nigrofimbriata ( Villeneuve, 1935 ) ♂ ­ Phallapodeme less arched; notch of syncercus funnel­shaped, without teeth ............. ............................................................................... * P. officialis ( Townsend, 1934 ) ♂