Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera
Author
Winterton, Shaun L.
Author
Martins, Caleb Califre
Author
Makarkin, Vladimir
Author
Ardila-Camacho, Adrian
Author
Wang, Yongjie
text
Zootaxa
2019
2019-04-09
4581
1
1
99
journal article
27348
10.11646/zootaxa.4581.1.1
5b259853-01f6-4307-8bef-58c98ec73bd3
1175-5326
2633842
20A9776D-AE5F-41BC-A35B-0C5E42EDFE48
Lithosmylidia
Riek, 1955
(
Fig. 5
)
Type
species.
Lithosmylidia lineata
Riek, 1955
: 678
(by original designation).
Diagnosis
. Forewing medium sized; trichosors present; crossveins poorly preserved in wings of all specimens; costal area with veinlets widely spaced, mostly simple; subcostal area with
sc-r
crossveins not detected; crossveins in area between RA and RP few or absent (one detected in one specimen of
L. parvula
); RP originating close to wing base, with at least 10 branches, very slightly sinuous distally in some species; crossveins of RP field not well preserved in most material, but apparently restricted to a few random ones proximally and two widely separate and irregular gradate series; M forked close to wing base; CuA irregularly pectinate forked, with branches oblique. CuP dichotomously forked with few branches; 1A pectinate, length variable; 2A deeply forked; 3A short and apparently simple; HW unknown.
Comments
. A specimen of
L. lineata
was considered by
Riek (1955)
as a hind wing, however
Lambkin (1988)
instead argued that it represented a forewing. Originally,
Lithosmylidia
was described by
Riek (1955)
belonging to
Kempyninae
, but
Makarkin
et al.
(2014)
argued that the genus belonged to
Archeosmylidae
. Characters such as Sc+RA joining the wing margin before the wing apex and the lack of RP crossveins support its placement in
Archeosmylidae
; unfortunately, important information about the hind wing cubital and anal vein fields were not preserved in any specimen.
Included species
.
L. baronne
Lambkin, 1988
(
Australia
)
(Middle Triassic)
L. lineata
Riek, 1955
(
Australia
)
(Late Triassic)
L. parvula
Riek, 1955
(
Australia
)
(Late Triassic)