European Rhogogaster s. str., with notes on several Asian species (Hymenoptera: Tenthredinidae)
Author
Taeger, Andreas
Author
Viitasaari, Matti
text
Zootaxa
2015
4013
3
369
398
journal article
10.11646/zootaxa.4013.3.3
fb7d2185-52ce-4846-8eb1-854c68574aab
1175-5326
245770
7C9F9BF9-2AC4-41B3-A2E6-2CE9F241C46A
Rhogogaster sibirica
Enslin, 1912
,
stat. nov.
(
Figs 1
e, 3j,
4m
,
6g
)
Rhogogaster viridis
var.
sibirica
Enslin, 1912
: 93
.
Lectotype
♀
hereby designated
(ZSM, http://dx.doi.org/10.6084/ m9.figshare.1335872).
Type
locality: “Irkutsk Sibir.”.
Paralectotypes
1♂
http://dx.doi.org/10.6084/m9.figshare.1117951, 1♀ http://dx.doi.org/10.6084/m9.figshare.1119167 (SDEI, see discussion).
Rhogogaster kudianus
(recte
kudiana
)
Rohwer, 1925
.
Holotype
♀ (USNM, http://usnmhymtypes.com/ default.asp?Action=Show_
Types
&Single_
Type
=True&TypeID=6086, not examined).
Type
locality: Kudia River, Amagu, Siberia (= Primorskiy Kray, Amgu River - A. Lelej, personal communication; ca.
45.83°N
,
137.67°E
),
syn. nov.
Discussion
. Describing
Tenthredo chlorosoma
,
Benson (1943)
treated
R. viridis
var.
sibirica
as a questionable synonym of
chlorosoma
. However, the examination of the
sibirica
types
showed, that these represent a species of the
viridis
group. The penis valves of the
paralectotype
(
Fig.
6g
) agree with the valves of
californica
(
Fig.
6
m).
Rhogogaster sibirica
might be an extremely pale form of
californica
, but currently no intermediate forms are known. Therefore it seems advisable to treat
R. sibirica
as a valid species.
The
type
status of the
sibirica
specimens in the SDEI requires an explanation: in 1910 Enslin published a revision of the Palearctic
Rhogogaster
. He described under
viridis
a very pale form from Siberia4, but gave no name for these specimens. His note “habe ich [...] Exemplare gesehen” (“I have seen specimens”) indicates that he had no
sibirica
material of his own.
In 1912
he slightly modified his 1910 description and named the form as
Rhogogaster viridis
var.
sibirica
.5 In the course of his
Rhogogaster
study of 1910, Enslin examined specimens from Konow’s collection, and in this collection (now in SDEI) is to be found a couple from “Irkutsk Sibir.” that fits Enslin’s description. These specimens have no identification labels by Enslin, but this is not surprising, because Enslin only later decided (1912) to give a name to this pale form. Enslin frequently kept dupla from Konow’s collection, and therefore it is to be expected that the specimen labeled “
Rhogogaster viridis var. sibirica Enslin
♀” (now coll. ZSM) is from the same series as the specimens in the Konow collection. Based on the discussion above, the specimen from the ZSM and the two specimens in the SDEI collection are considered to be
syntypes
.
Zirngiebl (1965)
noted that he has seen “den Typ”6 of
sibirica
from Irkutsk. This treatment does not fulfil the requirements of a
lectotype
designation in the ICZN Art. 74.5, as it is clear from the description that Enslin had
syntypes
before him, and the specimen was not explicitly selected from this series. The ♀ of the ZSM is hereby designated as
lectotype
.
Rhogogaster kudiana
Rohwer, 1925
, from the Russian Far East agrees with
sibirica
. A specimen from Anisimovka (Russian Far East, not far from the
type
locality of
kudiana
, http://dx.doi.org/10.6084/ m9.figshare.873867) was compared with the
sibirica
types
from Irkutsk, and also with the available figures of the
kudiana
type
. No relevant characters were found to distinguish two taxa. Our colleague D.R. Smith (Washington) kindly compared the
type
of
kudiana
with images of a specimen from Anisimovka, and provided some additional information about the
type
.
4.
Enslin (1910: 32, under
R. viridis
)
... “So habe ich namentlich aus Sibirien Exemplare gesehen, die auf den ersten Blick viel mehr der
Rh. punctulata
Kl.
gleichen, ja die schwarze Farbe ist bei ihnen noch geringer entwickelt. Es erscheint bei ihnen nur schwarz: am Kopfe ein feiner Saum um das untere Nebenauge und eine halbkreisförmige Linie, von den oberen Nebenaugen zu den seitlichen Stirnfurchen verlaufend; am Brustrücken nur teilweise die Nähte. Der Hinterleib ist durchaus grün, die schwarze Linie der Beine ist nur an den Tibien und den hintersten Schenkeln zu sehen. Tarsenspitzen und die meisten Adern und Nerven der Flügel sind schwarz.”
5.
Enslin (1912: 93, under
R. viridis
)
... “So ist bei der
var. nov
.
sibirica
der Körper fast ganz grün, nur eine wie bei
punctulata
ω-ähnliche Zeichnung des Oberkopfes—oft auch diese nur unvollständig—und am Thoraxrücken nur teilweise die Nähte schwarz, so daß das Tier heller gefärbt ist als
punctulata
; auch sind bei dieser
var. die
vorderen Schenkel auf der Hinterseite nicht oder nur an der Spitze schwarz liniert.”
6.
Zirngiebl (1965: 100)
“Den Typ der Enslinschen Form
var.
sibirica
habe ich gesehen. Mir diese Möglichkeit geboten zu haben, verdanke ich Herrn Dr. Franz Bachmair von den Zool. Sammlungen des Bayerischen Staates in München. Nach der Tabelle von Benson bestimmt, deckt sich das Tier mit der Form
chlorosoma
; es ist eher noch etwas heller. Es handelt sich um ein ♀, das so hell ist wie die ♂♂ und von Irkutsk aus Sibirien stammt.”