New Fossil Histerid Species from Cretaceous Burmese Amber (Coleoptera: Histeridae) Author Caterino, Michael S. text The Coleopterists Bulletin 2021 2021-03-20 75 1 211 221 http://dx.doi.org/10.1649/0010-065x-75.1.211 journal article 10.1649/0010-065X-75.1.211 1938-4394 5335054 urn:lsid:zoobank.org:pub:5C049771-EB80-4E0A-98AE-AD7CE31CAF57 Druantia Caterino , new genus zoobank.org/ urn:lsid:zoobank.org:act: E39E2917-9C2B-4936-8E4B-79BD0AC22FEE Type Species. Druantia aeterna Caterino , new species . Diagnosis. This extinct genus is distinguished by its rather elongate, subquadrate, slightly flattened body shape; frons with lateral carinae over antennal bases, extending to anterior margin of epistoma; labrum wide, short, plurisetose, but with prominent setae at each side; pronotum with posterior margin simply rounded; scutellar shield small if present (obscured by fracture); elytron without obvious dorsal striae, but weakly carinate outer marginal stria, apex weakly truncate; propygidium short, pygidium simple; procoxa strong, distinctly transverse; prosternal keel narrow, slightly produced posteriorly over anterior margin of mesoventrite; prosternal lobe short, incised laterally for passage of funicle, with anterolateral grooves for reception of major protibial spur; antennal cavities not well defined, open, antennal club apparently lying under the hypomeron in repose; mesoventrite not obviously emarginate, but overlain by apex of prosternal keel; metaventrite weakly convex; metanepisternum exposed, narrow; profemur with groove on inner surface to receive protibia; protibia with weakly defined depression for reception of tarsus on anterior surface, apex with strong outer spur directed laterad and smaller inner spur directed distad; tarsi 5-5-5 (probably—apical metatarsomeres missing); tarsal claws simple, separate. Derivation of Name. Druantia is based on the name of a fictional tree goddess, ruler of plant fertility, and protector of trees. Gender: feminine. Druantia aeterna Caterino , new species zoobank.org/ urn:lsid:zoobank.org:act: 0CBDFA69-956E-4C04-9BEC-8EF3CD225463 ( Figs. 14–17 ) Type Material. Holotype specimen, of unknown sex. Type locality: northern Myanmar : probably Hukawng Valley, collected during 2018; deposited in MSCC . The specimen was purchased by the author. The specimen bears a large fracture along and above the right elytron, obscuring that entire dorsal aspect. The amber in front of the head is slightly hazy. Numerous small air bubbles adhering to dorsal and ventral surfaces obscure details of surface sculpturing (in Figs. 16–17 ventral bubbles falsely appear to be punctures). Description. Total body (pronotum + elytra) length: 1.65 mm ; maximum (humeral) width: 0.94 mm ; pronotal length: 0.63 mm ; pronotal width: 0.88 mm . Body rather elongate, subquadrate, slightly flattened; frons convex; mandibles with long apices, left mandible apparently with tooth on incisor margin; antennal scape short, thick, funicle long, club tomentose, sutures or annuli unclear, if present; pronotum rather narrow, sides subparallel at base, rounded to weakly acute anterior corners, emarginate behind head; elytron without obvious dorsal striae, one epipleural stria and weakly carinate outer marginal stria (= outer subhumeral?) present; elytral apices weakly truncate; propygidium short, pygidium simple; procoxa strong, distinctly transverse; prosternal keel with a pair of striae along ridge, slightly produced over anterior margin of mesoventrite; prosternal lobe short; mesoventrite broad, with arched marginal stria; metaventrite weakly convex; profemur with groove on inner surface to receive protibia; protibia broad, outer edge rounded, bearing prominent spines and especially prominent apical corner, inner edge with distinct fringe of shorter setae; mesotibia rather slender, with ∼6 marginal spines and pair of small apical spurs; metatibia slightly broader, also with two small apical spurs; tarsi slender, apical tarsomere about 2.5× length of others, each with a pair of apicoventral spines. Derivation of Name. The species name is a Latin adjective and refers to the “eternal” preservation of this extinct species. Syninclusions. None. A DIAGNOSTIC CATALOG OF CRETACEOUS HISTERIDAE Antigracilinae † Antigracilus costatus Zhou, Caterino, Ren, and Ślipiński, 2020 Age: Barremian (125 Ma), compression fossil, Yixian Formation, China Diagnostic Features: Body loosely compact; frontoclypeal suture present; metacoxae narrowly separated. Abraeinae: Pantostictini † Pantostictus burmanicus Poinar and Brown, 2009 Age : Cenomanian (99 Ma), Burmese amber 218 THE COLEOPTERISTS BULLETIN 75(1), 2021 Figs. 14–17. Druantia aeterna , new species . 14) Anterior view; 15) Dorsal view; 16) Lateral view; 17) Ventral view. Diagnostic Features: Body elongate, narrowed anteriorly; dorsum simply punctate, lacking striae or carinae; ultimate maxillary palpomere large, fusiform; antennal cavity located at sides beneath pronotal hypomeron.