On the taxonomic status of Agelanius antenninus (Philip) (Diptera: Tabanidae) from Peru based upon evidence from a morphological analysis of the holotype Author González, Christian R. text Zootaxa 2023 2023-04-24 5271 1 163 168 http://dx.doi.org/10.11646/zootaxa.5271.1.7 journal article 10.11646/zootaxa.5271.1.7 7f72c2ad-4d8d-4cb7-9a82-13c8c68ed626 1175-5326 7864465 2C1C62CA-848D-4A74-80BD-01B9669777F7 Stypommisa ( Styphocera ) antennina Philip, 1969: 151–152 . ( Figs. 1A–F ) Stypommisa ( Styphocera ) antennina Philip, 1969 . Moucha, 1976: 232 . Dasybasis ( Agelanius ) antennina ( Philip, 1969 ) . Fairchild & Burger, 1994: 12 . Agelanius antennina ( Philip, 1969 ) . González, 1999: 143 . Agelanius antenninus ( Philip, 1969 ) . Coscarón & Papavero, 2009: 58 . Type material examined. Holotype , ♀ of Stypommisa ( Styphocera ) antennina PERU , Huánuco , Department Huánuco, deposited in the Cornell University Insect Collection ( CUIC ) No. 6747 ( CUIC 000058931 ), labeled as follows: “Huanuco, PERU, 17 Sept. 1954 , 7,500 ft” [white label with handwritten data]; “Holotype” “ Stypommisa ( Styphocera ) antennina ” n.sp. , Det. C.B. Philip 69 [red label with handwritten data] ( Fig. 1A ). The holotype is partly damaged. The head is detached, both basal flagellomeres are lost, and the one remaining wing is partially destroyed. The anterior legs are missing, and some of the legs lack tarsal segments. Redescription ( Holotype ♀ ) FIGURE 1. Holotype Stypommisa ( Styphocera ) antennina Philip, 1969 . A. Labels of holotype of Stypommisa ( Styphocera ) antennina Philip, 1969 . B. Head in frontal view. C. Head in ventral view. D. Thorax in dorsal view. E. Thorax in lateral view. F. Wing and abdomen. ©Cornell University Insect Collection. Diagnosis. Length 8.0 mm, wing length 8.5 mm . Body color dark brown. Eyes dark brown, with long, sparse ommatrichia. Frons moderately wide and divergent towards subcallus; frontal index 1.7. Basal callus dark brown, triangular with median dorsal projection reaching half the height of the frons. Ocellar triangle and ocelli vestigial. Scape of antenna subcylindrical, 2.5 times the length of pedicel; basal flagellomere indistinctly triannulated. Palpus slightly tapering distally and widening proximally, three-quarters the length of the proboscis. Mesonotum with dark brown integument, bearing sparse medium dark brown setulae. Wings slightly smoky, with dark brown clouds on the crossveins. Abdomen dark brown dorsally, distal margin of tergites lighter, sternites lighter yet. Head. Eyes dark brown with long, sparse ommatrichia. Frons moderately wide and divergent towards subcallus, with brownish pruinescence and with long brownish setulae ( Fig. 1B ); frontal index 1.7. Vertex with brownish pruinescence and long black setulae. Basal callus dark brown, triangular, with median dorsal projection reaching half the height of frons, touching subcallus but not touching eyes ( Fig. 1B ). Ocellar triangle and ocelli vestigial. Subcallus covered with brownish pruinescence, without setulae. Gena with brown pruinescence and with long brownish setulae. Clypeus with grayish pruinescence and with long brownish setulae, medially without setulae. Scape of antenna subcylindrical, 3.0 times length of pedicel, reddish brown, with long erect black setulae. Pedicel short, concolorous, bearing short black setulae, without dorsal projection ( Fig. 1C ). Basal flagellomere “robust not excised dorsally, style chunky and indistinctly 3-annulated.” Palpus with brownish pruinescence, with long black and grayish setulae, setulae longer on first segment than on second segment; first segment dark brown, one-quarter length of second segment; second segment lighter in color, slightly tapering distally and widening proximally, three-quarters length of proboscis, inner side without setulae ( Fig. 1C ). Proboscis with dark brownish-black setulae, labellum large and fleshy ( Fig. 1C ). TABLE 1. Morphological differences between Stypommisa ( Styphocera ) Philip and Agelanius Rondani.

Character	Stypommisa ( Styphocera ) Philip	Agelanius Rondani
Size	Small (8.00 mm)	Medium, large (12.5–15.5 mm)
Basal callus shape	Triangular (Fig. 1B)	Keel-shaped (Fig. 2A)
Frons width	Wide (Fig. 1B)	Narrow (Fig. 2A)
Frons lateral margins	Divergent (Fig. 1B)	Parallel (Fig. 2A)
Frontal index	1.7	2.5–3.4
Ommatrichia	Sparse	Abundant (Fig. 2B)
Ocellar triangle	Vestigial (Fig. 1B)	Well developed (Fig. 2C)
Ocelli	Vestigial (Fig. 1B)	Well developed (Fig. 2C)
Scape	Large, 3.0 times the length of pedicel	Medium, 2.0 times the length of pedicel (Fig. 2D)
Pedicel	Without dorsal projection (Fig. 1C)	With dorsal projection
Number apical flagellomeres	3	4 (Fig. 2D)
Palpus	Slightly tapering distally and widening proximally (Fig. 1C)	Slender and elongate (Fig. 2E)
Palpus setulae	Long	Short (Fig. 2E)
Scutum	Without narrow longitudinal stripes (Fig. 1D)	With narrow longitudinal stripes (Fig. 2F)
Vein R 4	With a short appendix	Appendix variable (Fig. 2G)
Abdomen	Without triangles (Fig. 1F)	Generally with triangles (Fig. 2F)
Distribution	Peru	Southern Argentina and Chile

Thorax. Scutum with dark brown integument, bearing sparse long dark brown setulae; postpronotal lobes light brown with long blackish setulae. Scutellum with long setulae laterally, disc with shorter setulae ( Fig. 1D ). Pleura covered with dark brown pruinescence, with long light brown setulae; anepisternum with long dark brown setulae; katepisternum and katatergite with long light brown setulae ( Fig. 1E ). Legs light brown, coxae darker, tibiae reddish brown, femora and tibiae with abundant long black setulae; remaining leg segments with short black setulae. Wing (partially destroyed). Wing membrane slightly infumated, with dark brown clouds on crossveins, cell r 5 widely open; veins Sc and R 1 with abundant short black setulae along their entire length; vein R 4 with short appendix ( Fig. 1F ). Basicosta with short black setae. Halter dark brown. Squamae concolorous with wing base. Upper calypter light brown, lower calypter lighter in color. Abdomen. Dark brown dorsally, distal margin of tergites lighter, tergites 2–7 with light brown setulae, longer setulae laterally. Sternites pale brown with long light brown setulae ( Fig. 1F ). FIGURE 2. Female of Agelanius meridianus Rondani. A. Head in frontal view (scale bar = 1 mm). B. Head in lateral view (scale bar = 0,5 mm). C. Vertex and ocellar triangle (scale bar = 0,5 mm). D. Antenna in lateral view (scale bar = 0,5 mm). E. Palpus in lateral view (scale bar = 0,5 mm). F. Habitus in dorsal view (scale bar = 1,0 mm). G. Wing (scale bar = 1,0). H. Labels of specimens. Male . Unknown. Distribution . Known only from the type locality