A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae) Author Means, Jackson C. https://orcid.org/0000-0001-7377-0696 Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA mjacks4@vt.edu Author Hennen, Derek A. https://orcid.org/0000-0001-7005-1151 Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA Author Marek, Paul E. https://orcid.org/0000-0002-7048-2514 Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA text ZooKeys 2021 2021-04-13 1030 1 180 http://dx.doi.org/10.3897/zookeys.1030.62544 journal article http://dx.doi.org/10.3897/zookeys.1030.62544 1313-2970-1030-1 875199397EEE5F7898EA1DB25DA62D25 Nannaria "Happy Valley" incertae sedis Fig. 119 Material examined. United States - North Carolina • 1 ♂ and 2 ♀♀ ; Caldwell County , 5 NW Lenoir , nr. Jct . US 321 & NC 268, on paved drive nr. woods; 35.9819°N , - 81.5641°W ; 19 Nov. 1979 ; E. Newton leg.; NCSM NAN0499. For detailed collection data see Suppl. material 7 . Hypothesized placement. Nannaria "Happy Valley" specimens are hypothesized to be closely related to Nannaria minor based on geographic proximity and the following combination of gonopodal characters: With wide distance between bases of acropodite and prefemoral process. Presence of medial flange near apex. Short, blunt distal zone. Nannaria "Happy Valley" specimens differ from N. minor based on the following combination of gonopodal characters: Acropodite medial flange much reduced in "Happy Valley". Prefemoral process directed ventrally at tip, not anteriorly as in N. minor . Medial curve of acropodite abrupt, forming a nearly 90° angle (Fig. 119A , red arrow), not smooth and gradual as in N. minor . Acropodite basal zone extremely wide in medial view (Fig. 119B , red triangle). Figure 119. Nannaria "Happy Valley" incertae sedis ♂ (NCSM, NAN0499) left gonopod A anterior view; red arrow indicates abrupt medial bend of acropodite B medial view; red triangle indicates wide basal zone C posterior view. Scale bar: 0.5 mm.