Systematics Of The Short-Tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), With Descriptions Of Six New Genera And Five New Species
Author
Monjaraz-Ruedas, Rodrigo
Author
Prendini, Lorenzo
Author
Francke, Oscar F.
text
Bulletin of the American Museum of Natural History
2019
2019-06-20
2019
435
1
93
https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2019/issue-435/0003-0090.435.1.1/Systematics-of-the-Short-Tailed-Whipscorpion-Genus-Stenochrus-Chamberlin-1922/10.1206/0003-0090.435.1.1.full
journal article
10.1206/0003-0090.435.1.1
0003-0090
5416046
Heteroschizomus goodnightorum
Rowland, 1973
Figures 4
, 6, 8E, 21A–C; table 7
Heteroschizomus goodnightorum
Rowland, 1973a: 2–6
;
1973b: 202
;
1973c: 136
;
1975b: 31
;
Rowland and Reddell, 1977: 83
;
Reddell and Cokendolpher, 1991: 18
;
1995: 3
, 5, 101, 106.
Schizomus goodnightorum
:
Rowland 1975a: 34
, 167, 168, 194, 257–269, 325, 397;
Rowland and Reddell, 1977: 90
, 99, 100; 1979a: 163, 164; 1981: 19, 21–27;
Reddell and Cokendolpher, 1995: 3
, 106.
Stenochrus goodnightorum
:
Reddell and Cokendolpher, 1991: 18
;
1995: 5
, 8, 18, 102, 106;
Villarreal et al., 2008: 64
;
Villarreal and García, 2012: 5
; Monjaraz-Ruedas and Francke, 2016: 783, 784, 797, 804; 2018: 190.
TYPE MATERIAL
:
MEXICO
:
Yucatan
: Chichén Itzá,
vi.1948
,
C
. Goodnight,
holotype
♂
,
paratype
♂
(
AMNH
).
SUPPLEMENTARY DESCRIPTION: The following description of the female is based on the additional material examined.
Prosoma
: Propeltidium with two apical setae; three pairs of dorsal setae.
Chelicerae
: Movable finger with very small lamella; serrula with 13 teeth, guard tooth present. Fixed finger with five small teeth between two primary teeth; setal group formula, 3-6-4-2-7-8-1-6.
Pedipalps
: Pedipalps homeomorphic; 1.59× longer than propeltidium. Trochanter with small rounded apical process, not projected; prolateral surface with small apical spur. Femur 1.73× longer than high; retroventral margin with setae
Fe
1
,
Fe
5
,
Fev
1
, and
Fev
2
acuminate; prolateral surface with row of three ventral setae (
Fmv
1–3
) and two dorsal setae (
Fmd
2
,
Fmd
3
). Patella with three acuminate
Pe
setae and four feathered
Pm
setae; without distinctive armature. Tibia setal formula, 3:3:4;
Ter
acuminate,
Tmr
and
Tir
feathered. Tarsal spurs asymmetric.
Opisthosoma:
Segments X–XII not elongated.
Pygidial flagellum
: Flagellum (
♀
) with three flagellomeres (fig. 21A–
C
); seta
Dl
2
reduced, situated posterior to
Vl
1
,
Dl
3
situated posterior to
Vl
2
,
Vm
2
present,
Vm
1
aligned with
Vm
2
;
Dl
1
and
Dl
4
microsetae present.
Female spermathecae
: Two pairs of lobes (fig. 8E); median and lateral lobes conical, subequal in length and width; median lobes linear, lateral lobes sublinear, apex slightly curved laterally, unsclerotized apically, with few duct openings; median lobe bases aligned with lateral lobe bases. Chitinized arch mug shaped; anterior branch not sclerotized. Gonopod wide; width ca. 2×length.
REMARKS:
Rowland and Reddell (1977)
synonymized
Heteroschizomus
,
stat. rev.
, and transferred
H. goodnightorum
to
Schizomus
, based on the discovery of other species with an elongated opisthosoma.
Reddell and Cokendolpher (1991)
transferred
Schizomus goodnightorum
to
Stenochrus
based on the morphology of the female spermathecae described by
Rowland and Reddell (1981)
. The discovery and correct identification of the female of
H. goodnightorum
confirmed the validity of
Heteroschizomus
and the correct placement of
H. goodnightorum
and related species, discoveries that were independently supported by the DNA sequence data.
ADDITIONAL
MATERIAL
EXAMINED:
MEX- ICO:
Yucatán
:
Municipio de Chichén Itzá
:
Chichen Itzá
,
vi.1948
,
C
.
Goodnight
,
1 ♀
(
AMNH
)
;
Fuera de la Cueva de Kanthon
,
20°38′08″N
88°29′42″W
,
23 m
,
2.viii.2014
,
J. Mendoza
and
R. Monjaraz
,
4 ♂
(AMCC [LP 14522]),
2 ♂
,
1 ♀
(CNAN Sz171).
Municipio
de Kaua
:
Fuera de la Cueva de Kaua
,
20°37′04″N
88°25′26″W
,
24 m
,
5.x.2014
,
G. Contreras
and
G. Montiel
,
1 ♂
,
2 ♀
(CNAN Sz163).
Quintana Roo
:
Municipio
de Cancun
:
Km
234
Highway
Mérida
–
Cancun
,
20°53′57″N
87°28′25″W
,
19 m
,
20. vii.2010
,
D. Barrales
,
G. Contreras
,
J. Cruz
,
O. Francke
,
G. Montiel
,
M. Paradiz
,
C
.
Santibañez
, and
A. Valdez
,
1 ♂
(CNAN Sz8)
.