A Taxonomic Revision Of The Spider Genus Meriola Banks (Araneae: Trachelidae) Author González, María E. 0000-0003-1609-0314 mariaegm86@gmail.com Author Grismado, Cristian J. 0000-0002-1260-7964 grismado@macn.gov.ar Author Ramírez, Martín J. 0000-0002-0358-0130 ramirez@macn.gov.ar text Zootaxa 2021 2021-02-24 4936 1 1 113 journal article 7437 10.11646/zootaxa.4936.1.1 5404afb9-168e-4d0f-b386-da1c314df1d9 1175-5326 4559165 DAC2F49B-EE13-4827-B549-C59B9C43550B Meriola cetiformis Strand Figs 1F , 19–21 , 90 Gayenna alticola Simon, 1896: 400 (male paralectotype , misidentified, from La Paz , Bolivia , in MNHN, examined by Ramírez, 2003 ); Simon, 1897: 94 , fig. 87. Trachelas cetiformis Strand, 1908: 241 (three female syntypes from Yauli, Junín , Peru , in MWG, examined by Platnick & Ewing, 1995 ). Trachelas 4-punctatus Mello-Leit„o, 1922: 49 (one male [lacking both palpi] and five female syntypes from Pinheiro , Rio de Janeiro , Brazil , formerly in MNRJ, probably lost; examined by Platnick & Ewing, 1995 ); Camargo , 1953: 324, figs 10–17. Trachelas caxambuensis Mello-Leit „o, 1926: 13, figs 14–15 (one male and one female [lacking abdomen] syntypes from Caxambui , Minas Gerais , Brazil , formerly in MNRJ, probably lost; examined by Platnick & Ewing, 1995 ). Trachelas distinctus Mello-Leit „o, 1941a: 178, fig. 69 (female holotype from Yala , Jujuy , Argentina , in MLP, examined). Cetonana cinerea Mello-Leit „o, 1941b: 221, fig. 32 (male holotype from Reconquista , Santa Fe , Argentina , in MLP, examined). Meriola arequipa Gertsch, 1942: 11 , fig. 34 (female holotype [lacking abdomen] from Arequipa , Arequipa , Peru , in AMNH, examined by Platnick & Ewing 1995 ). Cetonana lissopalpus Mello-Leit „o, 1943b: 116, fig. 17 (male holotype from Los Molles , Río Atuel , Mendoza , Argentina , in MLP, examined). Trachelas quadripunctatus Roewer, 1955: 588 . Trachelas arequipa Roewer, 1955: 587 . Meriola cetiformis Platnick & Ewing, 1995: 15 , figs 29–33. Diagnosis. Males ( Figs 20–21 ) can be easily recognized by the long embolus that coils around the palp, and the copulatory bulb that is half the size of the cymbium. Females ( Fig. 19 ) are easily recognizable by the twisted copulatory ducts and the rounded secondary spermathecae ( Platnick & Ewing 1995 : fig. 33). Description. See Platnick & Ewing (1995) for male and female descriptions. FIGURE 18. Meriola californica Banks , male (AMNH MGM-00487). a , habitus, anterior view; b , left palp, ventral view; c , same, prolateral view; d , same, retrolateral view. FIGURE 19. Meriola cetiformis Strand , female (MACN-Ar 32374). a , habitus, dorsal view; b , same, ventral view; c , same, lateral view; d, prosoma, dorsal view; e , same, ventral view; f , same, lateral view; g , habitus, anterior view; h , epigyne, ventral view. FIGURE 20. Meriola cetiformis Strand , male (MACN-Ar 32375). a , habitus, dorsal view; b , same, ventral view; c , same, lateral view; d , prosoma, dorsal view; e , same, ventral view; f , same, lateral view; g , female, live specimen; h , male, live specimen. FIGURE 21. Meriola cetiformis Strand , male (MACN-Ar 32375). a , habitus, anterior view; b , left palp, ventral view; c , same, prolateral view; d , same, retrolateral view. Natural history and habitat. The species is widely distributed over several ecoregions, from open forests with low trees, shrubs and other herbs (especially rich in grasses and legumes) to savannas and gallery forests, xeric deciduous forests with a layer of gramineous, cacti and land bromeliads, and also in urban habitats. Several specimens were collected on forests with tall trees, vines and epiphytes, woody bamboos and tree ferns. A few specimens were collected between 3000–5000 m altitude, where arborescent cacti, shrubs and grasses abound. Other specimens were collected on shrub steppes and grasslands. Distribution. Known from central and southern Perú , Bolivia , southern Brazil , Uruguay and Argentina ; in Chile known only from the far north. New records: PERÚ : Ancash : Callejon de Huaylas, S 09.166666° , W 77.750000° , elev. 3048–4572 m , VIII.1988 , V. & B. Roth, 1 female (CAS, CASENT 9059472). Huaraz, S 09.526223° , W 77.528630° , 6.XII.1980 , C. Gold, 1 female (CAS, CASENT 9059202). Parque Nacional Huascarán, Laguna Querococha, S 09.727222° , W 77.329722° , elev. 4024 m , 17.V.2010 , L. Figueroa, R. Pinto-da-Rocha & D. Silva, 2 females (MUSM-ENT 0503129). Junín : Ondores, S 11.066154° , W 76.158344° , elev. 4080 m , 19.III.2007 , on rocks, near lake, G. Alvares & W. Paredes, 2 females , 1 male (IBSP 73270). Yaulí, La Oroya, Quebrada Cerro Quinan (Tupac Amaru), S 11.552083° , W 75.931111° , elev. 4119 m , 22.VIII.2005 , W. Paredes, 1 male (MUSM-ENT 0504595). Lima : Chinchorros, S 12.062485° , W 77.103070° , 13.XII.1980 , C. Gold, 3 females (CAS, CASENT 9059158). Cuzco : nr. Huancarane, along road from Cusco to Shintuya, 1st truck breakdown, S 13.498798° , W 71.665990° , 23.IX.1987 , J. Coddington, 2 females (NMNH 1561). Sacsayhuamán, S 13.509844° , W 71.981681° , 12.XII.1980 , C. Gold, 1 female (CAS, CASENT 9059198). Puno : Amantaní, S 15.666208° , W 69.710821° , elev. 3900 m , III.1948 , F. Blancas, 2 females (MUSM-ENT 0504602). Chucuito, Ancomarca, S 16.648331° , W 69.364453° , elev. 4081 m , 4.III.2010 , E. Razuri, 1 female (MUSM-ENT 0501378). Arequipa : Cerca a Paty, S 16.086208° , W 70.966024° , 27.I.2000 , J. Ochoa, 1 female (MUSM-ENT 0504599). BOLIVIA : Murillo: La Paz : Laguna Valle de Zongo, Viscachavi, S 16.194638° , W 68.126772° , 5.VIII.1993 , H. Höfer, 1 female (MCN 24108). Chacaltaya, S 16.347150° , W 68.127841° , elev. 4500 m , 5.VII.1993 , H. Höfer & A. Brescovit, 1 female , 1 imm. (IBSP 11895), 1 female (MCN 24089). Viscachani, S 16.470998° , W 68.214424° , SMNK, 1 female (IBSP 11855). BRAZIL : Minas Gerais : Vale Rio Santo Inácio Coromandel, S 18.697424° , W 46.940677° , 2.V.2001 , R. Gallon, 1 female (IBSP 28558). Pudente de Morais: Fazenda do Sapé, rodovia MG-424, S 19.468383° , W 44.241606° , 2.XI.2000 , E. Álvares, 1 female (UFMG 318). Same locality, 3–4.II.2001 , E. Álvares, 1 female (UFMG 315). Viçosa, S 20.754888° , W 42.878610° , 2004, J. Lopes, 2 males (IBSP 55830). Coimbra, S 20.857272° , W 42.801276° , 5.II.1999 , pitfall, F. Marquini, 1 female (IBSP 23862). Espirito Santo : Bom Jesus do Itabapoana: Usina Hidrelétrica de Rosal, Rio Itabapuana, S„o José do Calçado, S 20.916943° , W 41.721666° , IX.1999 , I. Knysak, 2 females , 4 males (IBSP 26372). Mato Grosso do Sul : Anaurilândia: Usina Hidrelétrica Engenheiro Sérgio Motta, S 22.500000° , W 53.016667° , 15.XI–23.XII.1998 , Eq. IBSP, 2 females , 2 males (IBSP 23395). São Paulo : Usina Hidreléctrica Três Irm„os, entre Andradira y Barreto, S 20.672420° , W 51.299264° , IX.1990 , Costa & Bertim, 1 male (IBSP 4809).Jaboticabal, S21.252514° , W 48.325676° , 4.IX.1986 , H. Cunha, 3 females , 1 male (MCN 17809), 1 female (MCN 18785). Ibó, Rio Claro: Santa Rita do Passa Quatro, S 21.686507° , W 47.488274° , 3.XI.1940 , O. Schubart, 1 female (MZUSP 8542). Águas da Prata, S 21.917661° , W 46.685365° , IX.2004 , V. Onofrio, 1 male (IBSP 70982). Same locality, 21.I.1999 , V. Onofrio, 1 male (IBSP 22061). Anaurilândia: Usina Hidrelétrica Engenheiro Sérgio Motta, President Epitácio, S 22.500000° , W 53.016667° , 16.I–13.II.1999 , equipe IBSP, 1 female (IBSP 23048). Same locality, I–II.2000, equipe IBSP, 5 females, 2 males (IBSP 30106). Americana, S 22.737846° , W 47.333569° , 1944, R. Eller, 1 male , 2 imms. (MZUSP 5994). Guarulhos, S 23.454251° , W 46.533798° , 1.XI.1943 , F. Pereira, 1 female (MZUSP 12521). Baruerí, S 23.511369° , W 46.872942° , 26.XII.1965 , K. Lenko, 3 females (MZUSP 12522). Osasco, S 23.532857° , W 46.791952° , XI.1972 , A. de Souza, 1 female , 1 male (IBSP 3253). Perdizes, S 23.536902° , W 46.674317° , 12.IX.1951 , H. Urban, 1 male (MZUSP 1433). Río Claro: Colégio Claretiano, S 23.542080° , W 46.654604° , 17– 19.I.1962 , B. Reichardt & S. Penna, 1 female (MZUSP 11766). Capital, S 23.550520° , W 46.633309° , VIII.1941 , M. Maurus, 1 female (MZUSP 739). Butantã: Jardim Rizzo (URB.USP), S 23.572008° , W 46.732739° , 24.II.1999 , shrubs, D. Candiani, 1 female (IBSP 28819). Same locality, 1.III.1999 , M. Sebastiao (col. X IBMa herbaceo), 1 female (IBSP 28912). Ribeirão Preto: Ipiranga, S 23.585498° , W 46.599011° , 19.IV.1942 , J. Barroso, 1 female , 1 male (MZUSP 8368). Paraná : Flora da Serra, Boa Vista da Aparecida, S 25.436411° , W 53.407367° , 7–14.X.1998 , Eq. IBSP, 1 female (IBSP 21123). Dois Vizinhos: Foz do Chopim, Cruzeiro do Iguaçu, S 25.523865° , W 53.096533° , 8–15.X.1998 , equipe IBSP, 1 female (IBSP 21257). Santa Catarina : Estrada Nova Teutonia-Itá, S 27.183333° , W 52.483611° , 1.II.1996 , A. Bonaldo, A. Kury & R. Rocha, 1 female , 1 male (MCN 27133). Florianopolis: Ilha do Árvoredo, Reserva Biológica Marinha de Arvoredo, S 27.285247° , W 48.365877° , 2003, Noili, 1 male (IBSP 75355), 1 male (IBSP 75356). Rio Grande do Sul : Derrubadas: Parque Estadual do Turvo, S 27.206705° , W 53.935427° , 25–30.IV.2005 , R. Ott et al ., 1 male (MCN 39423). Aratiba: Usina Hidreléctrica de Itá, S 27.277222° , W 52.383333° , collector unknown, 1 female (MCTP). Represa de Garabí, S 28.020047° , W 55.225725° , collector unknown, 1 female (MCTP), 1 female (MCTP), 1 male (MCTP). Canela, S 29.356493° , W 50.812426° , 31.XII.1973 , A. Lise, 1 female (MCN 2064). Taquara, S 29.650973° , W 50.775785° , 18.VI.1988 , F. Quadros, 1 male (MCTP 26097). Capão da Canoa: Xangri-Lá, S 29.802808° , W 50.041231° , 9.XI.1986 , A. Lise, 1 female , 2 males (MCN 16007). Same locality, 20.II.1990 , A. Lise, 1 male (MCN 19569). Santo Antônio da Patrulha: Chicolom„, S 29.957592° , W 50.583635° , 23.III.1983 , H. Gastal, 1 female (MCN 11493). Porto Alegre, S 30.034656° , W 51.217658° , 13.VII.1962 , R. Krebs, 1 male (MCN 1279). Viamão: Aguas Belas, S 30.082400° , W 51.019946° , 24.I.1977 , A. Lise, 1 male (MCN 5157). Guaíba, S 30.114759° , W 51.322857° , 22.I.1975 , H. Gastal, 1 female (MCN 2771). Vila Nova, S 30.115413° , W 51.213207° , 23.VI.1991 , F. Garcia, 1 female (MCTP 2765). Serraria, Ipanema, S 30.150859° , W 51.214094° , 3.XI.1962 , C. Haktlieb, 1 female (MCN 1334). São Sepé, S 30.203212° , W 53.606907° , 25.VI.2000 , T. Gomes, 1 male (IBSP 122641). Capivari do Sul : Lagoa do Capivari, S 30.222000° , W 50.548400° , 20.V.2004 , Equipe Probio, 2 females (MCN 37379), 1 female (MCN 37394). Reserva Biológica do Lami José Lutzenberger, S 30.246942° , W 51.106937° , 15.VI.2000 , E. da Silva, 2 females (IBSP 48899), 1 female , 1 male (IBSP 48929). Same locality, 20.VIII.2000 , E. da Silva, 1 female (IBSP 48930). Quaraí: Estância S„o Roberto, S 30.293520° , W 55.961878° , 24–28.V.1991 , A. Brescovit, 1 female (MCN 21105). Palmares do Sul : Praia do Quintão, S 30.318719° , W 50.346460° , 10.XII.1988 , A. Bonaldo, 1 male (MCN 17968). Caçapava do Sul : Santa Barbinha, S 30.514853° , W 53.483217° , 30.VI.2005 , L. Almeido, 23 females , 10 males (MCTP 17214). Santana do Livramento: Estância Bela Vista, S 30.716799° , W 55.689897° , 26.V.2005 , J. Bitencourt, 1 male (IBSP 84903). Cerro Verde, S 30.805520° , W 55.531290° , 21–28.XI.2006 , J. Dias, 3 females , 2 males (IBSP 79920). Same locality, 28–29.XI.2003 , J. Bitencourt, 1 female , 1 male (IBSP 84964). Same locality, 3–9.XI.2006 , J. Dias, 4 females , 7 males (IBSP 79912). Same locality, 21–28.IX.2006 , J. Bitencourt, 1 female (IBSP 85002). Cerrito, S 31.689398° , W 52.831266° , 24.I.2008 , J. Rosado, 1 female (MCN 46720). Rio Grande, S 32.299886° , W 52.446547° , II.2005 , F. dos Santos, 1 female (MCTP 21794). Same locality, VIII.2004 , F. dos Santos, 2 females (MCTP 21792). Same locality, XI.2004 , F. dos Santos, 1 male (MCTP 21793). Santa Vitória do Palmar: Estaç„o Ecológica do Taim, S 32.491554° , W 52.584327° , 08.IV.1986 , A. Lise, 1 male (MCN 14857). Same locality, 9.IV.1986 , M. Marques, 1 female (MCN 14884). URUGUAY : Montevideo : Hill 500 m S of Castle, 15.XII.1965 , R. Capocasale & L. Bruno, 1 female (CAS, CASENT 9059407). Colonia : Colonia del Sacramento , S 34.460719° , W 57.833910° , 11.XII.2005 , G. Ruiz & C. Rheims, 2 females , 2 males (IBSP 58680). ARGENTINA : Salta : Anta: Parque Nacional El Rey, S 24.684121° , W 64.638642° , I.1981 , P. Goloboff, 1 female , 1 male (MACN-Ar 29744). Misiones : Iguazú: Parque Nacional Iguazú, S 25.68029° , W 54.319289° , I.1993 , C. Grismado, 1 male (MACN-Ar 16359). Tucumán : El Infiernillo (AOP-loc-002), S 26.74272° , W 65.77283° , 18.X.2014 , under rocks at the edge of a cliff, A. Porta, 1 female, sample MGM-00422 (MACN-Ar 33018). Tafí del Valle, Cerro El Pelao (AOP-loc-01), S 26.87206° , W 65.73646° , 17.X.2014 , under rocks at the edge of a cliff, A. Porta, 1 female , sample MGM-00419 (MACN-Ar 33015), 1 female , sample MGM-00420 (MACN-Ar 33016), 1 male , sample MGM-00421 (MACN-Ar 33017). Tafí del Valle: Parque Los Menhires, S 26.930705° , W 65.680000° , 4.VIII.1990 , A. Brescovit & A. Bonaldo, 1 female , 1 male (MCN 19945). Corrientes : Mburucuyá: Parque Nacional Mburucuyá, Camino del Uno, S 28.000999° , W 58.095255° , elev. 64 m , 27–30.V.2011 , G. Rubio, L. Piacentini & M. Izquierdo, 1 female , 1 male (MACN-Ar 29743). Bella Vista, S 28.507731° , W 59.044857° , 7.X.2005 , G. Ávalos, 1 male (CARTROUNNE 8270). La Rioja : Famatina: Cueva de Noroña, S 28.091944° , W 67.673056° , elev. 2846 m , 29.I.2006 , L. Piacentini, L. Compagnucci & A. Ojanguren Affilastro, 8 females , 2 males , 2 imms. (MACN-Ar 35193). Santa Fe : General Obligado: 3 km NE de Berna (MJR-loc-165), S 29.26211° , W 59.81745° , elev. 43 m , 19–24.III.2014 , dry chacoan forest with sandy soil, hand collection, M. Ramírez, C. Grismado, L. Piacentini & M. González Márquez, 1 female , sample MGM-00273 (MACN-Ar 33010). 9 de Julio: Ruta Provincial Nº 13, 2,45 km S de cruce con Ruta Nacional Nº 98, 81,3 km por aire W de Vera (MJR-loc-162), S 29.28352° , W 61.027° , elev. 70 m , 19–24.III.2014 , flooded grasslands, hand collection, M. Ramírez, C. Grismado, L. Piacentini & M. González Márquez, 1 female , 1 imm. (MACN-Ar 33009). General Obligado: Berna, Arroyo El Palmar, S 29.2842222° , W 59.8613333° , elev. 47 m , 5–7.VI.2011 , grassland, hand collection, G. Rubio, L. Piacentini & M. Izquierdo, 1 female (MACN-Ar 29741). Vera: Las Gamas, bañado a la entrada de la estancia, S 29.41233° , W 60.37423° , elev. 61 m , 5–7.VI.2011 , rushes, hand collection, G. Rubio, M. Izquierdo, L. Piacentini & R. Adilardi, 2 males (MACN-Ar 29742). Estancia Las Gamas, bañado sobre Ruta Nacional Nº 98, 16,5 km por aire W de Vera (MJR-loc-160), S 29.41382° , W 60.37438° , elev. 23 m , 19–24.III.2014 , marsh, hand collection, M. Ramírez, C. Grismado, L. Piacentini & M. González Márquez, 1 female , sample MAI- 04176, MGM-00357 (MACN-Ar 32374), 1 male , temporary preparations MGM-00358–00359 (MACN-Ar 32375), 1 male (MACN-Ar 33011). La Capital: San José del Rincón, S 31.602076° , W 60.577061° , XII.1995 , C. Scioscia, 1 female (MACN-Ar 16368). San Juan : Iglesia: Ruta Nacional Nº150, camino a Paso de Agua Negra, 6 km W de Arrequintín, S 30.369475° , W 69.65721° , 23.I.1982 , E. Maury, 2 females , 1 male (MACN-Ar 16406). Paso de Agua Negra, S 30.39291° , W 69.566809° , III.2006 , A. Ojanguren Affilastro & L. Compagnucci, 1 male (MACN-Ar 35164). Calingasta: Parque Nacional El Leoncito, S 31.863213° , W 69.226267° , 25.I.2006 , L. Piacentini, L. Compagnucci &A. Ojanguren Affilastro, 1 female (MACN-Ar 12859). Córdoba: Capital: Barrio Argüello, S 31.348071° , W 64.25982° , 12.X.2002 , C. Mattoni, 1 female , 1 male (MACN-Ar 31202). Entre Ríos : Colón: Parque Nacional El Palmar (MJR-loc-17), S 31.86534° , W 58.23745° , elev. 22 m , 6.VIII.2011 , rocks, palm trees with grassland and lowland forest, M. Ramírez & Equipo MACN-Ar, 1 female , sample MGM-00318 (MACN-Ar 32987). Parque Nacional El Palmar, Sector Sur (MJR-loc-22), S 31.88768° , W 58.31186° , elev. 30 m , 7.VIII.2011 , grassland and lowland forest with palm trees (bosque bajo), M. Ramírez & Equipo MACN-Ar, 1 female (MACN-Ar 32775). Diamante: Parque Nacional Predelta, Campo Sarmiento, pastizal, S 32.100711° , W 60.643125° , elev. 35 m , 23– 26.V.2011 , grassland, hand collection, G. Rubio, L. Piacentini & M. Izquierdo, 1 female , 1 imm. (MACN-Ar 29739). Parque Nacional Predelta, isla frente a seccional, 6 km S de Diamante (MJR-loc-143), S 32.1214° , W 60.63528° , elev. 10 m , 1.V.2013 , forest of Tessaria integrifolia (“aliso”), Mikania cordifolia , Cayaponia citrullifolia and Ephedra tweediana (“enredaderas”), beating shrubs, M. Ramírez, L. Piacentini, M. González Márquez, A. Laborda & S. Aisen, 1 male , sample MGM-00143 (MACN-Ar 30281), 1 female , sample MGM-00144 (MACN-Ar 30282). Parque Nacional Predelta, bañado al pie de bosque de barranca, S 32.12154° , W 60.62946° , elev. 4 m , 23–26.V.2011 , rushes, hand collection, G. Rubio, L. Piacentini & M. Izquierdo, 1 female (MACN-Ar 29738), 2 females , 1 male (MACN-Ar 30990). Parque Nacional Predelta, Campo Sarmiento, 6 km S de Diamante (MJR-loc-141), S 32.12182° , W 60.62525° , elev. 39 m , 30.VI.2013 , grassland at the edge of Baccharis sp. vegetation (“Chilcal”), hand collection, M. Ramírez, L. Piacentini, M. González Márquez, A. Laborda & S. Aisen, 1 female , 1 male (MACN-Ar 30829). Parque Nacional Predelta, bañado cerca de entrada, 6 km S de Diamante (MJR-loc-142), S 32.12194° , W 60.62996° , elev. 10 m , 30.VI.2013 , marsh in lagoon, hand collection, M. Ramírez, L. Piacentini, M. González Márquez, A. Laborda & S. Aisen, 1 female , sample MGM-00131 (MACN-Ar 30269), 1 female , sample MGM-00138 (MACN-Ar 30276). Parque Nacional Predelta, sendero a Laguna Las Piedras, 6 km S de Diamante (MJR-loc-140), S 32.12438° , W 60.63002° , elev. 6 m , 29.V–1.VI.2013 , forest of Tessaria integrifolia (“aliso”), Mikania cordifolia , Cayaponia citrullifolia and Ephedra tweediana (“enredaderas”), beating shrubs, M. Ramírez, L. Piacentini, M. González Márquez, A. Laborda & S. Aisen, 1 male , sample MGM-00150 (MACN-Ar 30288). Parque Nacional Predelta, sendero a Laguna Las Piedras, S 32.12462° , W 60.6277° , elev. 12 m , 23–26.V.2011 , hand collection, G. Rubio, L. Piacentini & M. Izquierdo, 1 female (MACN-Ar 29737), 1 male (MACN-Ar 29740). Ciudad Autónoma de Buenos Aires : Avenida Warnes 42, S 34.604169° , W 58.437973° , 28.III.2013 , M. Ramírez, 1 female , sample MGM-00129 (MACN-Ar 30267). Buenos Aires : S 36.412991° , W 60.402024° , VI.1939 , B. Gerschman de Pikelin & R. Schiapelli, 1 female (MACN-Ar 618). San Pedro: Reserva Natural Histórica y Refugio de Vida Silvestre Municipal Vuelta de Obligado, S 33.583333° , W 59.816667° , elev. 14 m , 9.I.2009 , buffer zone, hand collection, N. López Carrión & N. Olejnik, 1 male (MACN-Ar 36969). Same data, 10.IV.2009 , 1 female , 2 males (MACN-Ar 36877), 1 male (MACN-Ar 36969), 3 males (MACN-Ar 36997). Same data, 23.V.2009 , 3 males (MACN-Ar 36883). Same date, 24.V.2009 , 10 females , 2 males (MACN-Ar 36884). Same data, 18.VII.2009 , 2 females (MACN-Ar 36887). Same data, 11.X.2009 , 1 male (MACN-Ar 36954). Delta del Río Paraná: 3° Sección San Fernando: Río Barca Grande y A° Barquita, S 34.119725° , W 58.444922° , 24–26.I.2012 , C. Grismado & L. Zapata, 1 male (MACN-Ar 30981). Isla Martín García, S 34.182558° , W 58.251352° , I.1940 , J. Viana, 2 females , 1 male (MACN-Ar 16622). Campana: Reserva Natural Otamendi, S 34.230186° , W 58.869773° , 7.VI.2007 , hand collection, C. Grismado, L. Damer, N. López, S. Trivero, I. Crudele & N. Olejnik, 1 male (MACN-Ar 12422). San Antonio de Areco, S 34.255921° , W 59.467433° , VI.1982 , M. Ramírez, 1 female (MACN-Ar 30980). Campana: Estación Río Luján, S 34.278° , W 58.890168° , 7.VI.2007 , grassland, beating, C. Grismado, L. Damer, N. López, S. Trivero, I. Crudele & N. Olejnik, 1 female , 1 male (MACN-Ar 12381). Same locality, 9.VI.2007 , in nest of Metazygia gregalis , hand collection, C. Grismado, 1 female (MACN-Ar 12520). Same locality, 30.VI.2007 , grassland, beating, C. Grismado, L. Damer, N. López, S. Trivero & N. Olejnik, 2 females , 3 imms. (MACN-Ar 12689), 1 female , 3 imms. (MACN-Ar 12739). Delta del Río Paraná, Delta del Tigre, S 34.319621° , W 58.628883° , X.1940 , J. Viana, 1 male (MACN-Ar 16624). Tigre: reserva hotel “Amarran Sancho”. Pajonal [LNP-loc-45c], S 34.36377° , W 58.59774° , 11–12.VI.2015 , beating shrubs, C. Grismado, L. Piacentini, N. Lopez Carrión, D. Proud & J. Varano, 1 female (MACN-Ar 34578). Ferrocarril General Belgrano, Campo de Mayo, km 26, S 34.556174° , W 58.657914° , 5.II.2006 , nest of “leñatero” ( Anumbius annumbi ), P. Turienzo, 1 female , 5 imms. (MACN-Ar 33084). Tres de Febrero: Caseros, S 34.610233° , W 58.567302° , 11.XII.1946 , collector unknown, 1 female (MACN-Ar 16372). Florencio Varela, S 34.795673° , W 58.2706° , II.1939 , F. Monrós, 1 male (MACN-Ar 16623). Ensenada: Punta Lara, S 34.805302° , W 58.033479° , 11.X.1981 , F. Miranda & M. Ramírez, 1 female (MACN-Ar 30978). Spegazzini: Paraná de las Palmas, S 34.883358° , W 58.637966° , 20.VI.1963 , M. Galiano, 1 female (MACN-Ar 30982). Magdalena, S 35.07892° , W 57.51798° , 7.II.1982 , P. Goloboff & M. Ramírez, 1 male (MACN-Ar 30977). Punta Indio: Punta Piedras, S 35.420706° , W 57.133876° , 13.III.1983 , P. Goloboff & E. Maury, 1 female , 1 male (MACN-Ar 16345). Carlos Casares, S 35.626417° , W 61.357787° , 1979, P. Goloboff, 1 female (MACN-Ar 16343), 1 male (MACN-Ar 16376). Chascomús: Paraje El Destino, Ruta Provincial Nº57 entre Lezama y Pila, banquina este [MJR-loc-184d], S 35.943639° , W 57.996776° , elev. 13 m , 20.VIII.2015 , grassland (“Pastizal pampeano”), hand collection, M. Ramírez, C. Grismado, L. Piacentini, M. Izquierdo, S. Aisen, D. Proud, N.L. Carrión, I. Magalhaes & R. Botero, 6 females , 2 males , 1 imm. (MACN-Ar 33096), 23 females , 9 males (MACN-Ar 33097), 49 females , 35 males , 52 imms. (MACN-Ar 33098), 17 females , 6 males , 6 imms. (MACN-Ar 33099), 78 females , 31 males , 1 imm. (MACN-Ar 37407). Las Flores: Rosas, Estación del Ferrocarril General Roca, S 35.966782° , W 58.939918° , collector unknown, 1 male (MACN-Ar 16625). General Madariaga, S 37.170088° , W 57.317254° , I.1962 , M. Galiano, 1 female (MACN-Ar 5364). Tandil, S 37.31566° , W 59.142152° , I.1982 , M. Ramírez, 1 male , 1 imm. (MACN-Ar 30979). Same locality, 18.XII.1997 , C. Grismado, 1 male (MACN-Ar 16355). Neuquén : Añelo: Punta Carranza, S 37.973258° , W 69.148505° , 23.XI.1973 , O. de Ferraris, 1 male (MACN-Ar 30565).