Revision and phylogenetic analysis of Chilicola sensu stricto (Hymenoptera: Colletidae) with the description of a new species Author Gibbs, Jason Author Packer, Laurence text Zootaxa 2006 1355 1 37 journal article 10.5281/zenodo.174590 0d36725c-b244-4aeb-a0fa-d1cd09dbb987 1175-5326 174590 Subgenus Chilicola Spinola s. str. Chilicola Spinola, 1851 , Hist. Fis. Pol. Chile . Zool. Type Species: C. rubriventris Spinola, 1851 , designated by Sandhouse, 1943 Diagnosis. The following unique characteristics allow species of the subgenus Chilicola s. str. to be differentiated from related subgenera: face slightly concave in lateral view [ Fig. 14I ; Chilicola (Chilioediscelis) has erroneously been reported to also have a concave face ( Michener 1995 , 2000 )], distal margin of clypeus curved around lateral margin of labrum ( Fig. 14B ), S6 of females with apical spine ( Fig. 15 L), mesepisternum and S2 of males with dense pubescence ( Figs. 12D, 12H respectively), S7 of males with curved, apically oriented dorsal process ( Figs. 1–6 , 16B ), Chilicola s. str. and C. (Chilioediscelis) share the following: vertex sometimes concave ( Fig. 14 K), hind tibial spurs robust and curved ( Fig. 15F ), frons of females with longitudinal striae ( Fig. 13D ), fore femur of males robust, and spiculum moderately broad and long ( Fig. 16D ). Chilicola s. str. displays the following plesiomorphies that differentiate it from C. (Chilioediscelis) : male clypeus with granulose sculpture ( Fig. 12K ), pronotal lobe entirely black ( Fig. 10A ), episternal groove extends below scrobal groove ( Fig. 13H ), inner tooth of hind claw well developed ( Fig. 15G ), and S2 scopa of females corbiculate ( Fig. 12I ). Description. Black-brown; males with variable patterns of yellow on face and legs except anterior surface of male fore tibia always yellow ( Figs. 10E ) and fore and mid distitarsi always yellow-orange; females often with orange-red on metasoma ( Fig. 11H ); face slightly concave; distal margin of clypeus curved around lateral margin of labrum; head broader than long; eyes convergent below; clypeus broader than long; frons with slight depression above antennal sockets; episternal groove extends below scrobal groove; hind femur of males robust with broad concavity on ventral surface continuous with flattened ventral surface of hind trochanter; hind tibial spurs robust and curved; hind tibia of some males expanded with ventral concavity ( Figs. 1E , 2C , 4E , 5C ); S1 of males with or without ventrally oriented process ( Figs. 15I , J ); males with dense hairs on S2; females with weak hind leg scopa; S2 of females with corbiculate scopa; microsculpture granular; punctation variable except dense on lower and upper paraocular areas and frons; S6 of females with apical spine; S7 of males with two lateral lobes, dorsal lobes curved, apically oriented ( Figs. 1–6 ); S8 with long, moderately broad spiculum; penis valve with pair of large, dorsal, membranous appendages ( Figs. 1–6 ).