Description of the nymph of Miroculis (Miroculis) eldorado Gama-Neto & Hamada, 2014 (Leptophlebiidae: Ephemeroptera) from Roraima, northern Brazil
Author
Boldrini, Rafael
Author
Lima, Lucas Ramos Costa
Author
Ottoni-Boldrini, Bianca Maira De Paiva
text
Zootaxa
2018
2018-08-14
4459
1
197
200
journal article
29037
10.11646/zootaxa.4459.1.12
f475bef2-1873-4aaa-947a-a21397d0a9ed
1175-5326
1458332
E9E6BDD9-274C-4BCD-B678-76D9AB261456
Miroculis
(
M
.)
eldorado
(
Figs. 1
–10)
Diagnoses.
Male imago:
1) forewing with membrane hyaline, distal area of costal and subcostal region lightly darker; 2) eleven facets in the longest row of upper portion of compound eye; 3) femora of all legs with an apical blackish band, hind femur with a medial blackish band; 4) penes long and thin, wider basally, length equal to segments I+II of forceps.
Nymph
: 1) anterolateral margin of labrum slightly flattened with lateral margins angular (
Fig. 4
); 2) Segment II of labial palp 1.15 x length of Segment I (
Fig. 8
); 3) Segment II of maxillary palp 1.34 x the length of Segment I (
Fig. 7
); 4) posterolateral spines on terga VIII–IX larger (
Fig. 3
); 5) gills with lateral lobes well-developed and trachea unbranched (Fig. 10).
Mature nymph.
Lengths. Body: 5.0–
5.5 mm
. Foreleg:
2.88 mm
; middle leg:
2.72 mm
; hind leg: 3.08 (all coxa and trochanter not included). Caudal filament: broken.
Coloration
(
Figs. 1–3
).
Head:
brown with pale area surrounding ocelli and eyes; lateral edges of genae darker; antenna with scape and flagellum pale yellow and pedicel darker. Eye: upper portion of turbinate eye reddish brown and lower portion black; ocelli apically purplish and basally blackish. Mouthparts: mandibles dark brown; maxillae pale yellow; hypopharynx pale yellow; labium light brown; submentum light brown.
Thorax:
pro-, meso-, and metanotum brown with medial pale yellow line; antelateroparapsidal sutures and sublateroscutum blackish (
Fig. 1
). Thoracic sterna whitish, except metasternum with anterior margin dark brown.
Legs:
pale yellow; femora with sub-apical blackish bands (
Figs. 1, 2
); ventral margin of tibia with blackish mark and subapical blackish band.
Abdomen:
abdominal terga I–X brown with a pair of medial macula and lateral blackish marks; tergum X darker; sterna pale yellow (
Fig. 1
). Gills gray and trachea blackish gray (Fig. 10). Caudal filaments pale yellow; basal portion of joint darker.
Morphology,
Head:
clypeus concave anteriorly; labrum with anterolateral margin slightly flattened and lateral margins angular (
Fig. 4
); outer margin of mandibles curved (
Figs. 5, 6
); maxilla as in
Fig. 7
; inner margin of apex on SII of labial palp not acute (
Fig. 8
). Width of Labium palp Segment I 0.58 x length; Segment III length of labial palp 0.53 x length of Segment II and 0.61 x length of Segment I; Segment II length 1.15 x length of Segment I. Maxilla: Segment III of maxillary palp 0.44 x length of Segment II and 0.59 x length of Segment I; Segment II 1.34 x length of Segment I.
Legs:
Foreleg: width of femur 0.31 x length; width of tarsal claw 0.33 x length; midleg: width of femur 0.32 x length; hind leg: width of femur 0.31 x length. subapical denticle of tarsal claw of foreleg larger than the remaining denticles (
Fig. 9
).
Abdomen:
posterolateral spines on segments IV–IX (
Fig. 3
) (larger on segments VIII–IX). Gill: Length of medial filament 0.98 x length of body of gill; trachea unbranched, lateral lobes present, well-developed; inner lobe of dorsal lamella 3.7 x length of outer margin (Fig. 10).
Life cycle association.
Male imago reared from nymph.
Distribution.
Roraima
State,
Brazil
.
Material
examined.
One
nymph,
Brazil
,
Roraima
,
Amajari
,
Serra do Tepequém
,
Igarapé Geladeira
,
03°47'N
/
61°45'W
,
01.xi.2014
,
Boldrini, R.
and
Barroso, P.C.S.
col. Two nymphs,
Brazil
,
Roraima
,
Amajari
,
Serra do Tepequém
,
Igarapé do Barata
,
03°47'07.7"N
/
61°45'02.6"W
,
16.ii.2016
,
Boldrini
, R. col.
One
male imago (reared, exuviae mounted on slides),
Brazil
,
Roraima
,
Amajari
,
Serra do Tepequém
,
Igarapé do Barata
,
03°47'07.7"N
/
61°45'02.6"W
,
01.xii.2017
, Boldrini, R. and Lima, L.R.C. col.
Comments.
The nymph of
Miroculis
(
M.
)
eldorado
, shares some similar characters with
Miroculis
(
Atroari
)
amazonicus
Savage & Peters
and
M.
(
Miroculis
)
fittkaui
Savage & Peters
, especially by having gills with grayish coloration and trachea unbranched, posterolateral lobes well-developed, and labrum with lateral margins angular (
Fig. 4
). However,
M.
(
M.
)
eldorado
can be differentiated from both by the posterolateral abdominal spines: on segments IV–IX in
M.
(
M.
)
eldorado
, II–IX in
Miroculis
(
Atroari
)
amazonicus
and
M
. (
M
.)
fittkaui
.