Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models Author Casali, Daniel M Author Boscaini, Alberto Author Gaudin, Timothy J Author Perini, Fernando A text Zoological Journal of the Linnean Society 2022 2022-12-01 196 4 1505 1551 https://academic.oup.com/zoolinnean/article/196/4/1505/6617197 journal article 10.1093/zoolinnean/zlac041 0024-4082 7381236 Nothrotheriinae plus Megatheriinae PP = 54, age = 17.96 Mya (16.39–19.87). This clade, which unites the clades Nothrotheriinae and Megatheriinae , was recovered only for H models and par_IW10 and par IW5, as stated above, and was poorly supported. The clade was supported by four synapomorphies, recovered with both methods: presence of lingual and labial grooves in mf3; temporal lines approximate midline but do not meet to form a sagittal crest on skull roof; well-developed buccinator fossa of maxilla, with a deep depression; and Glaserian fissure opening into a distinct groove in squamosal lying medial to entoglenoid process.