A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804
Author
Marusik, Yuri M.
Author
Omelko, Mikhail M.
text
Zootaxa
2014
3894
1
journal volume
10.11646/zootaxa.3894.1.4
1bff28d0-db79-4a0f-8cfa-fc6cd82ce2ae
1175-5326
251800
70B9B045-9C14-47B1-B0C7-8DDA09AFE02B
Gnaphosa koponeni
sp. n.
Figs 1–9
, Map 1.
Material examined
.
Holotype
♂ (
ISEA
),
Russia
, SE Tuva, East of Tannu-Ola Mt. Range,
20 km
of Khol’-Oozhu Vill.,
50°50’N
94°19’E
, mountain
Larix
-moss-stony forest-tundra,
2175 m
, 8–
16.06.1995
(D.V. Logunov).
Paratypes
:
1♂
2♀ (
ISEA
), together with
holotype
;
4♂
(
ZMUT
), Tuva, Tannu-Ola, S slope
50°50’N
,
94°18’E
,
2120 m
,
Pinus
cembra-Larix
forest, 8–
17.06.1995
(S. Koponen);
2♂
(
ZMUT
), West Sayan, Oiskiy Pass,
1650 m
, alpine tundra slope,
21.06.1995
(S. Koponen).
Etymology
. The species name is a patronym honouring its collector, our friend and colleague Seppo Koponen, University of Turku,
Finland
.
FIGURES 1–6.
Male palp of
Gnaphosa koponeni
sp. n.
1–2 palp, ventral and retrolateral; 3–6 bulbus, prolateral, ventral, from above and retrolateral. Scale = 0.2 mm. Abbreviations:
Bm—
bulging margin of cymbium,
Ea—
angle of embolus,
Eb—
base of embolus,
Tt—
strong teeth of embolus.
Diagnosis
. Males of the new species are most similar to
G. wiehlei
. The two species can be distinguished by a long tibial apophysis, large, widely spaced teeth at base of the embolus and distinct tapering of the terminal half of embolus (in prolateral view) in the new species (shorter tibial apophysis, one strong tooth (spur) on prolateral part of embolus, and gradual tapering of embolus in
G. w i e h l e i
). Males of the two sibling species can be separated on the basis of tibia I spination: the new species has only 2 apical spines, while
G. wiehlei
has 2 prolateral, 2 pairs of ventral and 2 apical spines (
Fig. 21
). Females of the new species are most similar to
G. sticta
in well developed bulges of the fovea, but can be distinguished by the wider scape in the new species, which is larger than the fovea bulge (smaller than the bulge in
G. s t i c t a
), larger glands of receptacles bent posteriorly (shorter and horizontally spread glands in
G. sticta
) and shape of lateral foveal margins: without pocket in new species and distinct pocket in
G. sticta
.
Females of
G. koponeni
sp. n.
differ from
G. wiehlei
by undeveloped bulges in
G. wiehlei
, short lateral foveal margins without pocket (long lateral margins with distinct pocket in
G. wiehlei
). Both sexes of the new species differ from
G. wiehlei
by having dark-brown coloration (light yellow in
G. wiehlei
).
FIGURES 7–15.
Epigynes of
Gnaphosa koponeni
sp. n.
(7–9),
G. wiehlei
(10–12) and
G. sticta
(13–15, from Finland). 7, 10, 13 intact, ventral; 8, 11, 14 macerated, ventral; 9, 12, 15 macerated, dorsal. Scale = 0.2 mm. Abbreviations:
Fb—
bulge of fovea,
Id—
insemination duct,
Lm—
lateral margin,
Mp—
pocket of lateral margin,
Pf—
posterior furrow,
Ps—
pocket of scape,
Re—
receptacle,
Rg—
gland of receptacle.
Description
. Male. Total length 6.35. Carapace 3.1 long, 2.4 wide. Carapace dark brown without pattern, fovea black. Sternum, labium, maxillae dark brown. Abdomen dark brown. Legs brown without spots. Legs spination: femora
I 2
d, 1p;
II 2
d, 2r;
III 2
d, 2p, 2r;
IV 2
d, 2p, 2r; patellae:
III 1
r, other without spines; tibiae
I 2
apical;
II 2
apical;
III 1
d, 4p, 3r, 2-
1v
and 2 apical;
IV 1
d, 3p, 4r, 2-
2v
and 2 apical; metatarsi I 1-
1v
; II 1-
1v
;
III 4
p, 4r, 2-0 (2-1)v and 4 apical;
IV 3
p, 4r, 2-
1v
and 6 apical. Length of legs joints as shown in
Table 1
.
TABLE 1.
Length of leg joints in male.
| Femur |
Patella |
Tibia |
Metatarsus |
Tarsus |
Total |
| I |
2.2 |
1.2 |
1.55 |
1.4 |
1.15 |
7.5 |
| II |
1.95 |
0.9 |
1.55 |
1.2 |
1.05 |
6.65 |
| III |
1.75 |
0.9 |
1.35 |
1.45 |
1.05 |
6.5 |
| IV |
2.2 |
1.2 |
1.85 |
2.25 |
1.25 |
8.75 |
Palp as in
Figs 1–6
; tibial apophysis as long as tibia, sharply pointed, undivided; retrobasal margin of cymbium bulging (convex) (
Bm
); median apophysis widened; embolus long, with strong teeth (
Tt
) on retrolateral part and angle (
Ea
) in mid part, after which the embolus tapers.
Female. Total length 9.9. Carapace 3.75 long, 2.7 wide. Coloration as in males. Legs spination: femora
I 2
d, 1p;
II 2
d, 1p;
III 2
(1)d. 2p, 2r;
IV 2
d, 1p, 1r; patellae:
III 1
r, other without spines; tibiae
II 1
apical;
III 1
d, 4p, 3r, 2-
2v
and 2 apical;
IV 4
p, 4r, 2-
2v
and 2 apical; metatarsi I 1-
1v
; II 1-
1v
;
III 4
p, 3r, 2-
1v
and 4 apical;
IV 3
p, 4r, 2-
2v
and 6 apical. Length of legs joints as shown in
Table 2
.
TABLE 2.
Length of leg joints in female.
| Femur |
Patella |
Tibia |
Metatarsus |
Tarsus |
Total |
| I |
2.4 |
1.3 |
1.75 |
1.25 |
1.1 |
7.8 |
| II |
2.1 |
1.2 |
1.6 |
1.25 |
1 |
7.15 |
| III |
1.75 |
1 |
1.3 |
1.55 |
1.05 |
6.65 |
| IV |
2.55 |
1.1 |
1.85 |
2.35 |
1.3 |
9.15 |
Epigyne as in
Figs 7–9
; scape large, wider than long, with concave posterior margin, pocket of scape (
Ps
) longer than wide; fovea with pair of large bulges (
Fb
) delimited by posterior furrows (
Pf
); lateral margins (
Lm
) short with almost undeveloped pocket (
Mp
); receptacles (
Re
) stretched almost horizontally; glands (
Rg
) relatively long, bent posteriorly.
Distribution
. The new species is known from the south part of the Krasnoyarsk Province and Tuva. Its occurrence in adjacent
Mongolia
is very likely. Some of the female records of
G. sticta
from Tuva may refer to this species.