Givetian ostracods of the Candás Formation (Asturias, North-western Spain): taxonomy, stratigraphy, palaeoecology, relationship to global events and palaeogeographical implications Author Maillet, Sebastien Author Milhau, Bruno Author Vreulx, Michel Author Posada, Luis-Carlos Sánchez De text Zootaxa 2016 4068 1 1 78 journal article 10.11646/zootaxa.4068.1.1 b9c1e3a3-e591-4b34-9f30-d6d88a230673 1175-5326 270644 80E89736-33F2-4FF5-9B01-8B7239407875 Cavellina cf. rhenana Krömmelbein, 1954 (Pl. II: Figs. 7 & 8) cf. 1954a Cavellina rhenana nov. sp.; Krömmelbein: 250–251, pl. 1, fig. 3. cf. 1983a Sulcella rhenana (Krömmelbein) ; Milhau: 351, pl. 1, figs. 7–8. cf. 1985 Cavellina rhenana Krömmelbein ; Coen: 18, pl. 5, figs. 5–8. cf. 1985 Sulcella rhenana (Krömmelbein) ; Lethiers et al. : 66–67, pl. 9, figs. 6–7. cf. 1988 Sulcella rhenana (Krömmelbein) ; Milhau: 487, pl. 56, fig. 13. cf. 2009 Cavellina rhenana Krömmelbein ; Casier & Préat: 96, pl. 5, fig. 3. non 2011b Cavellina cf. rhenana Krömmelbein ; Casier in Casier et al. : 111–113, pl. 2, fig. 13. cf. 2013 Cavellina rhenana Krömmelbein ; Casier & Maillet in Casier et al. : 264, fig. 9B. 2013 Cavellina cf. rhenana Krömmelbein ; Maillet: 85, pl. X, figs. 18–19. cf. 2013b Cavellina rhenana Krömmelbein ; Maillet et al. : fig. 5K. cf. 2013c Cavellina rhenana Krömmelbein ; Maillet et al. : figs. 2 A–K. PLATE II. Ostracods from the type-area of the carbonate Candás Formation. For samples stratigraphic position, see Figure 3. 1 –2 . Evlanella peranensis Maillet n. sp. , tecnomorph (1), N2, LV (C. O .F.L.S. 1126); heteromorph (2), N2, slightly oblique dorsal view (C. O .F.L.S. 1127); 3. Poloniella cf. claviformis ( Kummerow, 1953 ) , I6 , RV (C. O .F.L.S. 806); 4–5. Punctoprimitia cf. europaea Weyant, 1967 , adult (4), II1 , LV (C. O .F.L.S. 808); juvenile (5), II1 , LV (C. O .F.L.S. 809); 6. Cavellina ? wahlensis Coen, 1985 , tecnomorph, G1, LV (C. O .F.L.S. 812); 7–8. Cavellina cf. rhenana Krömmelbein, 1954 , heteromorph (7), M2, LV (C. O .F.L.S. 817); tecnomorph (8), M2, LV (C. O .F.L.S. 816); 9–10. Uchtovia refrathensis (Krömmelbein, 1954) , tecnomorph (9), M2, LV (C. O .F.L.S. 827); heteromorph (10), M2, LV (C. O .F.L.S. 1128); 11. Knoxiella ? sp. P1, I6 , LV (C. O .F.L.S. 830); 12. Cryptophyllus sp. P1, II1 , LV? (C. O .F.L.S. 857); 13. Cryptophyllus sp. P2, O 5, RV? (C. O .F.L.S. 858); 14. Cryptophyllus sp. P3, Q8, internal mould, LV? (C. O .F.L.S. 860); 15. Cryptophyllus sp. P4, M3, LV? (C. O .F.L.S. 838); 16. Roundyella ? sp. indet., I3 , LV? (C. O .F.L.S. 786); 17. Cytherellina aff. brassicalis Becker, 1965 , I6, RV (C. O .F.L.S. 872); 18. Cytherellina cf. dubia ( Kummerow, 1953 ) , G1, RV (C. O .F.L.S. 873); 19. Cytherellina cf. groosae ( Coen, 1985 ) , B10, RV (C. O .F.L.S. 874); 20. Cytherellina cf. perlonga ( Kummerow, 1953 ) , B10, RV (C. O .F.L.S. 876); 21. Cytherellina ? sp. P1, A9, RV (C. O .F.L.S. 881); 22. Cytherellina sp. indet., Q8, RV (C. O .F.L.S. 882); 23. Punctomosea ? sp. indet., Q8, RV (C. O .F.L.S. 893); 24. Polyzygia beckmanni beckmanni Krömmelbein, 1954 , V14, RV (C. O .F.L.S. 899); 25. Polyzygia insculpta insculpta Becker, 1964 , II1, RV (C. O .F.L.S. 901); 26–27. Polyzygia neodevonica neodevonica ( Matern, 1929 ) , adult (26), Q8, RV (C. O .F.L.S. 902); juvenile (27), M2, LV (C. O .F.L.S. 903); 28a, b. Polyzygia symmetrica symmetrica Gürich, 1896 , Q8, RV (a) and dorsal view (b) (C. O .F.L.S. 907). Material. 3 specimens (3 C). Description. Medium-sized sub-oval carapace, stocky in tecnomorphs and more elongated in heteromorphs. The larger RV overlaps the LV on whole outline, slightly anteriorly and more clearly ventrally, dorsally and posteriorly. Hmax at 1/3 posterior in tecnomorphs, more anterior in heteromorphs; Lmax slightly ventral; Wmax at 1/3 posterior. Lmax equivalent to about 2 Hmax. Slightly convex dorsal margin, anteriorly oblique (also posteriorly oblique in heteromorphs). Straight to slightly concave ventral margin. Dorsal and ventral margins almost parallel. Regularly convex, well rounded anterior margin. Well rounded posterior margin. Posterior part of the carapace higher than the anterior one. Surface slightly reticulated. Slightly anterior adductorial sulcus, more or less marked. Discussion. These specimens show a carapace with features close to C. rhenana Krömmelbein, 1954 , except that this one is much more elongate. A marked sexual dimorphism is observed, as usual in cavellinids. Not enough specimens were collected to observe the specific variation as known in C. rhenana ( Maillet et al. 2013c ). Occurrence. Middle Givetian (Candás Fm, mb C), Asturias (Peran-Perlora section).