Resurrection of the enigmatic species Prangos akymatodes (Apiaceae) from Central Afghanistan
Author
Lyskov, Dmitry
Department of Higher Plants, Faculty of Biology, Lomonosov Moscow State University, 1 - 12 Leninskie Gory, 119234, Moscow, Russia
Author
Samigullin, Tahir
Department of Evolutionary Biochemistry, Belozersky Institute of Physico-Chemical Biology, Lomonosov Moscow State University, 1 - 40 Leninskie Gory, 119234, Moscow, Russia
text
Phytotaxa
2017
2017-10-27
326
3
202
210
http://dx.doi.org/10.11646/phytotaxa.326.3.4
journal article
10.11646/phytotaxa.326.3.4
1179-3163
13698903
Prangos akymatodes
Rechinger & Riedl
in
Køie & Rechinger (1963: 111)
Type
:—
AFGHANISTAN
.
Shanbashak Pass
, Prov. Darra-i-Suf,
9000 ft.
, dry rocky slopes,
31.08.1939
(
31 Aug. 1939
),
Koelz
13875
(
lectotype
: US catalog No.: 3151765, barcode 00289235 [image!], designated here by Lyskov;
isolectotype
:
W
! barcode
W 1988- 0010066
)
.
Polycarpic, herbaceous plants with thick tap-root (
Fig. 3
). Stems
60–80 cm
tall and
5–6 mm
Ø
at base, mainly single, thin-grooved, scabrous, alternately branched from middle part, covered with remains of petioles and sheaths. Basal leaves formed rosette, petioles up to
8 cm
long, scabrous; leaf blades
18–22 cm
long,
10–15 cm
wide, 3–4 times pinnate; leaf segments from first to second with petiolules; terminal lobes
3–7 mm
long,
0.3–0.5 mm
wide, subulate, acute; stem leaves significantly smaller than basal leaves. Terminal umbels
8–10 cm
Ø
, with 13–16 rays; rays
4–5 cm
long, thickened, glabrous, unequal; umbellets up to
2.5 cm
Ø
, 10–15 flowers, with 2–4 linear, prickly, entire, bracteoles. Pedicels equal,
0.8–1.2 cm
long, terete, glabrous. Mericarps broadly elliptic (
Fig. 1A
), 10–15 ×
7–8 mm
, homomorphic, dorsally compressed; only primary ribs presented, all ribs straight, suberose, with continuous margin, lateral ribs more wide and well-developed; commissure intermediate; mesocarp divided into an outer “epimesocarp” and inner mesocarp; inner mesocarp divided into five completely separated parts, consisting of parenchymatous cells with thickened and pore walls, “epimesocarp” consisting of parenchymatous, nonlignified cells; endosperm with mushroom-like groove at commissural side.
Distribution:—
The species is known only from the
locus classicus
:
Afghanistan
, Darah Sof district, Shanbashak Pass.