The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Janssen, Ronald 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2022 2022-03-30 5123 1 1 172 http://dx.doi.org/10.11646/zootaxa.5123.1.1 journal article 10.11646/zootaxa.5123.1.1 1175-5326 6399827 036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A Neoperrona harzhauseri Kovács & Vicián, 2021 nov. comb. Figs 30A 1 –A 2 , B 1 –B 3 , C 1 –C , 4G , 5 , 7 Clavatula ( Clavatula ) cf. taurinensis May.— Pavlovsky 1957: 53 , pl. 2, figs 3a–b [ non Neoperrona taurinensis ( Bellardi, 1877 ) ]. * Perrona harzhauseri nov. sp. – Kovács & Vicián 2021: 146 , pl. 3, figs 21–29. Type material. Holotype : HNHM , PAL 2021.24 .1, SL: 54.0 mm, MD : 16.0 mm, Letkés ( Hungary ), illustrated in Kovács & Vicián (2021 , pl. 3, figs 21–22), figs 30A 1 –A 2 . Paratypes : NHMW , 2021/0002/0005, SL: 52.0 mm , MD : 16.6 mm , illustrated in Kovács & Vicián (2021 , pl. 3, fig. 23); NHMW , 2021/0002/0006, SL: 50.0 mm, MD : 15.5 mm , illustrated in Kovács & Vicián (2021 , pl. 3, fig. 26); HNHM , PAL 2021.25 .1, SL: 50.0 mm, MD : 16.0 mm, illustrated in Kovács & Vicián (2021 , pl. 3, figs 27–28); NHMW , 2021/0002/0007, SL: 51.0 mm, MD : 15.0 mm, illustrated in Kovács & Vicián (2021 , pl. 3, fig. 29); HNHM , PAL 2021.26 .1 illustrated in Kovács & Vicián (2021 , pl. 3, fig. 25); NHMW , 2021/0002/0008, illustrated in Kovács & Vicián (2021 , pl. 3, fig. 24); HNHM , PAL 2021.27 .1., PAL 2021.28 .1., PAL 2021.29 .1, all Letkés ( Hungary ). Illustrated material. NHMW 2019/0151/019a, SL: 49.2 mm , MD : 14.0 mm, Letkés ( Hungary ), figs 30B 1 –B 3 , 4G; NHMW 2019/0151/019, SL: 53.0 mm, MD : 16.2 mm , Letkés ( Hungary ), figs 30C 1 –C 2 . Additional studied material. NHMW 2019 /0151/019, 5 spec. , Letkés ( Hungary ) . Revised description. Shell moderately large, slender fusiform with very high spire; apical angle ~30°. Protoconch of two smooth convex whorls. Teleoconch of about 11 whorls. Early teleoconch whorls flat-sided, with tripartite sculpture; weak subsutural spiral cord, beaded suprasutural cord forming periphery, separated by concavity filled by smooth, flattened central cord. Mid-spire subsutural cord becomes swollen to form broad, smooth, sharply delimited collar, central cord remains narrow, suprasutural beads weaken leaving broad, smooth to vaguely tubercular rounded cord above suture. Suture indistinct on early whorls, narrowly impressed, linear on later whorls. Last whorl about 55% of total height; with broad subsutural collar. Subsutural ramp not well-defined, with smooth, narrow, low, rounded cord. Shoulder not developed, evenly rounded at periphery, moderately strongly constricted at base. Siphonal fasciole weakly swollen, narrow. Sculpture of irregular spirals below shoulder, most strongly developed at mid-whorl and over fasciole, weaker over base. Aperture moderately narrow, ovoid. Outer lip thin, smooth within. Anal sinus moderately, narrow, asymmetrically U-shaped, with apex at center cord. Siphonal canal moderately short. Columellar and parietal callus thickened, sharply delimited, forming moderately broad callus rim. Discussion. Perrona villarrasensis Vera-Peláez & Lozano-Francisco, 2001 , from the Tortonian and early Pliocene of Spain (= Pleurotoma pretiosa sensu Pereira Da Costa, 1867 , pl. 27, figs 1a–b) is superficially similar to Neoperrona harzhauseri Kovács & Vicián, 2021 , but is less slender, lacks a central spiral cord and has slightly more concave whorls (see Vera-Peláez & Lozano-Francisco 2001 , pl. 7, figs 1–6, pl. 9, figs 7–8; Landau et al. 2011 , pl. 21, figs 1–2). The same features separate Perrona cf. villarrasensis of Spadini & Manganelli (2010) , from the early Pliocene of Italy , from Neoperrona harzhauseri . We have not seen early teleoconch whorls of Perrona villarrasensis . Based on its general morphology, we assume that it is a Perrona . Paleoenvironment. Inner neritic environments with corals ( Kovács & Vicián 2013 ). Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin : Letkés ( Hungary ) ( Kovács & Vicián 2021 ); Pannonian Basin : Zapresic-Brijeg at Samobor ( Croatia ) ( Pavlovsky 1957 ).