New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia) Author Poletaev, Vladyslav 4BF22B2D-FD42-4588-AB80-BBA5DDF4A579 Institute of Geological Sciences of the National Academy of Sciences of Ukraine, 55 - b, Oles Honchar St., Kyiv, 01054, Ukraine. vipoletaev38@gmail.com text European Journal of Taxonomy 2024 2024-11-04 968 132 155 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2723/12515 journal article 305035 10.5852/ejt.2024.968.2723 21b497f6-a2ee-4af8-b2e0-027aadacb4eb 2118-9773 14047314 35322F0D-0A5C-4227-AB4D-D4758ECA6E7A Tiramnia davidi sp. nov. urn:lsid:zoobank.org:act: 0C805CE0-5009-4C43-BF07-087B0BBEB785 Fig. 3 Martinia ( Pseudomartinia ) sp. A. – Aisenverg 1950: 137 , pl. 2 figs 9–10. Martinia ( Pseudomartinia ) sp. – Aisenverg 1951: 43 , pl. 11 fig. 2. Martinia balkhashica – Poletaev 2008: 28 , pl. 1 figs 1–5. Tiramnia sp. – Poletaev 2018: 23 , pl. 13 figs 6–10. non Martinia balkhashica – Beznosova 1968: 191 , pl. 29 fig. 15. Diagnosis Medium-sized shell with angular edges of trough-shaped anterior part of sulcus of the ventral valve and a shallow furrow along axis of fold distinctly limited in the anterior part of the dorsal valve. Etymology Named in honor of David E. Aisenverg, who first distinguished this species. Material examined Holotype ( Fig. 3A ) UKRAINE • Donetsk Region , Khartsis’k District , Vovcha River , Illinka village ; late Bashkirian Smolyanynivka Formation , Н 4 limestone layer ; Aisenverg 1951 Coll.; NMNH 1433/85 . Paratypes ( Fig. 3B–E ) UKRAINE • 12 specs ; Luhansk Region , Bila River , Gorodyshche village ; late Bashkirian, Smolyanynivka Formation ; stratigraphic interval between the H 3 limestone layer to the H 6 limestone layer; Poletaev 2018 Coll.; NMNH 2446/102 to 2446/113 . Description Medium-sized shell ( 24 mm wide, 23 mm long and 17 mm thick in the holotype ), rounded pentagonal in outline and rather strongly and uniformly biconvex in lateral profile; hinge line shorter than the largest width at mid shell. Umbonal part of ventral valve broad, well-developed, with steep lateral slopes. Cardinal extremities rounded. Beak is well developed, inclined, projecting above hinge line. Ventral interarea short, narrow, triangle, smooth. Sulcus distinctly developed, rather wide, shallow with weak axial groove and trough-like tongue; uniplicate anterior margin with distinctly angular margins of tongue at junction with lateral slopes. Dorsal valve somewhat less convex than ventral one. Dorsal umbo small, well expressed. Fold broad, indistinct in posterior part, but distinctly developted in the anterior part, where its slopes fall steeply. Fold also bears a slight median furrow. Shell surface mostly smooth, weak lines of growth are visible, and rare traces of short weak radial ribs near the anterior commissure. Microsculpture not observed on the specimens. Internal structure of the ventral valve with subrhombic deepened muscular imprints in the umbonal region, as well as traces of single axial and several ramiform lateral imprints of the vascular system (see Fig. 3E ). Fig. 3. Tiramnia davidi sp. nov. from the Donets Basin (Ukraine). Pennsylvanian, Bashkirian. A . Holotype NMNH 1433/85 ( Аisenverg 1951 Coll.) Donetsk Region, Illinka village; ventral, dorsal, lateral and anterior views (copy from Аisenverg 1951 : pl. 11 fig. 2). B–C . Paratypes NMNH 2446/102, NMNH 2446/103 ( Poletaev 2018 Coll.) Luhansk Region, Bila River, Gorodyshche village; ventral views (copy from Poletaev 2018 : pl. 13 figs 8–9). D . Paratype NMNH 2446/104 ( Poletaev 2018 Coll.) same locality; dorsal view (copy from Poletaev 2018 : pl. 13 fig. 7). E . Paratype NMNH 2446/105 ( Poletaev 2018 Coll.) same locality; internal view of the ventral valve (copy from Poletaev 2018 : pl. 13 fig. 10). Scale bar units = 1 mm. Remarks The new species differs from other known species of Tiramnia in having a more pronounced sulcus with a trough-like profile of the tongue and a distinct fold with a median groove. Based on morphological similarity and close geological age, the specimens under study were previously assigned to the Bashkirian Martinia balkhashica Beznosova, 1968 from eastern Kazakhstan ( Poletaev 2008 ). However, the holotype of M. balkhashica has an interior with twin vascula media ( Beznosova 1968 : pl. 29 fig. 16), which clearly shows that the Kazakh species belongs to Jilinmartinia . Other representatives of Tiramnia , similar to T. semiglobosa , differ from the new species by the rounded contours of the shells, sometimes drop-shaped, with an elongated ventral apex, a weak, poorly defined sulcus, often with a median groove. Martinia buckmani Yanishevsky, 1918 from the late Serpukhovian of Central Asia and early Bashkirian of the Donets Basin and the Urals is very similar to the new species. However, M. buckmani is distinguished by its smaller size, swollen shell with a strongly protruding and wrapped apical part of the ventral valve, and apparently by its internal structure. Tiramnia davidi sp. nov. differs from Postmartinia juresanensis ( Stepanov, 1948 ) from the Pennsylvanian of Bashkortostan by the angular rather than rounded lateral profile of the clearly limited anterior end of the sulcus and its much less developed continuation (tongue), as well as by its stratigraphic and geographic occurrence.