The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition
Author
Harzhauser, Mathias
0000-0002-4471-6655
Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;
mathias.harzhauser@nhm-wien.ac.at
Author
Landau, Bernard
0000-0002-4471-6655
Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;
mathias.harzhauser@nhm-wien.ac.at
text
Zootaxa
2023
2023-11-14
5370
1
1
74
https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270
journal article
10.11646/zootaxa.5370.1.1
1175-5334
10147814
48903495-7C6C-46E4-9B1B-D6A2F2781873
Pseudotorinia grasemanni
nov. sp.
Figs 4N
,
19A–D
Architectonica
(
Nipteraxis
)
marthae
Boettger
—
Csepreghy-Meznerics 1956: 386
, pl. 1, figs 19–21 [
non
Pseudotorinia marthae
(
Boettger, 1902
)
].
Architectonica
(
Nipteraxis
)
martae
[sic] (Boettger)—Kojumdgieva in
Kojumdgieva & Strachimirov 1960: 93
, pl. 29, figs 17a–d [
non
Pseudotorinia marthae
(
Boettger, 1902
)
].
Solarium marthae
Boettger
—Strausz 1962: 122, pl. 51, figs 5–7 [
non
Pseudotorinia marthae
(
Boettger, 1902
)
].
Architectonica
(
Nipteraxia
[sic])
marthae
Boettger, 1901
—Strausz 1966: 118, pl. 51, figs 5–7 [
non
Pseudotorinia marthae
(
Boettger, 1902
)
].
Solarium marthae
Boettger
—
Csepreghy-Meznerics 1969b: 20
, pl. 2, figs 19, 23, 26 [
non
Pseudotorinia marthae
(
Boettger, 1902
)
].
Type material.
Holotype
:
NHMW 2023
/0267/0007, SL:
2.8 mm
, MD:
6.1 mm
,
Steinebrunn
(
Austria
),
Figs 19B
1
– B
3
.
Paratypes
:
NHMW 2023
/0267/0008, SL:
2.7 mm
, MD: 7.0 mm,
Steinebrunn
(
Austria
),
Figs 19A
1
–A
3
.
NHMW 2023
/0267/0009, SL:
2.5 mm
, MD: 6.0 mm,
Steinebrunn
(
Austria
),
Figs 19C
1
–C
3
.
NHMW 2023
/0267/0010, SL:
2.9 mm
, MD:
5.8 mm
,
Steinebrunn
(
Austria
),
Figs 4N
,
19D
1
–D
3
.
FIGURE 19 A
1
–A
3
.
Pseudotorinia grasemanni
nov. sp.
, paratype, NHMW 2023/0267/0008, Steinebrunn (Austria).
B
1
–B
3
.
Pseudotorinia grasemanni
nov. sp.
, holotype, NHMW 2023/0267/0001, Steinebrunn (Austria).
C
1
–C
3
.
Pseudotorinia grasemanni
nov. sp.
, paratype, NHMW 2023/0267/0009, Steinebrunn (Austria).
D
1
–D
3
.
Pseudotorinia grasemanni
nov. sp.
, paratype, NHMW 2023/0267/0010, Steinebrunn (Austria).
Type
locality.
Steinebrunn
(
Austria
),
Vienna
Basin
.
Type
stratum.
Silty sand of the Baden Formation.
Age.
Middle Miocene, early/middle Badenian (Langhian).
Etymology.
In honor of Bernhard Grasemann, Geologist at the Department of Geology of the University of
Vienna
(
Austria
).
Diagnosis.
Small, depressed lentiform shell with low conical to nearly flat spire and angled periphery with rounded keel. Dorsal sculpture of finely beaded spiral cords with SSC bearing slightly more prominent beads. Base with wide umbilicus and succession of five delicate beaded spiral threads on outer basal field and three stronger beaded spiral cords on the inner basal field.
Description.
Small, depressed lentiform shell with low conical to nearly flat spire. Protoconch slightly submerged below apex, small, of <1 visible whorl (diameter
0.6 mm
). Teleoconch of ~3.5 whorls. First teleoconch whorl flat with slightly raised, beaded SSC and UPC with two finer beaded MCs. Later whorls with very prominent SSC bearing rounded beads, three weaker MCs and prominent UPC. LPC separated from UPC by narrow but slightly deeper groove; largely covered by subsequent whorl causing relatively deep suture. Beads densely spaced, axially elongated, arranged in prosocline lines. LPC forming blunt keel; below steeply inclined nearly flat to faintly concave passing via weak angulation into weakly convex base. Periphery finely beaded in basal view. No IPC developed. Five narrow, densely beaded spiral cords on outer basal field and three broader beaded cords on inner basal field (innermost corresponds to PUC). Growth lines forming weakly cancellate pattern in interspaces between spiral cords. UC delimited from basal field by distinct groove, prominent, slightly sunken into and overhanging umbilicus. Umbilicus very wide (~31–38 of MD). Umbilical wall bearing fine axial growth lines. Aperture small, circular, CG and PG very weak.
Discussion.
Pseudotorinia grasemanni
is reminiscent of
P. marthae
(
Boettger, 1902
)
with which it has been confused in the literature. They can generally be separated by the wider umbilicus of
P. grasemanni
(~31–38 vs. ~20–26 of MD). Moreover, the beads on the periphery are coarser in
P. marthae
. There are several species described from the European Neogene, which agree with
Pseudotorinia grasemanni
in the wide umbilicus. Of these, the Pliocene to extant
Pseudotorinia architae
(O.G.
Costa, 1841
), from the Mediterraean Sea, might be a closely related species, but is distinguished by its comparatively uniform sculpture on the dorsal side and the rounded periphery, which is in a more adapical position (
Bieler 1993
: fig. 225;
Chirli 2013
; pl. 6, figs 3–9).
Pseudotorinia ivolasi
(
Ivolas & Peyrot, 1900
)
, from the Langhian of the
Loire
Basin (
France
), differs in its nearly flat spire and rounded periphery (
Glibert 1949
pl. 6, figs 15a-b, pl. 7, fig. 8).
Pseudotorinia planulatum
(
Grateloup, 1832
)
, from the Burdigalian of the
Aquitaine
Basin (
France
), differs in its axial sculpture on the basal field and the reticulate sculpture on the dorsum (
Cossmann & Peyrot 1919
: pl. 15, figs 71–73; https://science.mnhn.fr/institution/mnhn/collection/f/ item/j05829?listIndex=68&listCount=169).
Pseudotorinia subplicata
(d’Orbigny, 1852)
, from the Burdigalian of the
Aquitaine
Basin (
France
), differs in its rounded periphery (
Cossmann & Peyrot 1919
: pl. 15, figs 59–61). The Pliocene Mediterranean
Pseudotorinia obtusa
(
Bronn, 1831
)
differs in its narrower umbilicus, more rounded periphery, and more elevated spire (
Ferrero Mortara
et al.
1984
pl. 15, figs 4a–c).
Distribution.
Known so far only from the middle Badenian (late Langhian) of the Central Paratethys Sea.
Central Paratethys Sea.
Middle Miocene (Badenian):
Vienna
Basin
: Baden, Steinebrunn (
Austria
) (
hoc opus
); Pannonian Basin: Szob (
Hungary
) (Strausz 1966); B̧kk Mountains: Sámsonháza (
Hungary
) (
Csepreghy-Meznerics 1969b
);
Dacian Basin
: Opanec (
Bulgaria
) (Kojumdgieva in
Kojumdgieva & Strachimirov 1960
).