Discovery of Griphophanes Grootaert & Meuffels and Nepalomyia Hollis in the Afrotropical Region with a key to Afrotropical genera of Peloropeodinae (Diptera: Dolichopodidae) Author Ya, Igor text Zootaxa 2010 2668 1 20 journal article 10.5281/zenodo.276399 f90a978c-086a-4bf6-b8d7-575fabb91ef3 1175-5326 276399 Nepalomyia kotrbae sp. nov. ( Figs. 15−23 ) Diagnosis. Having simple male wing veins and simple hypandrium, short postpedicel of male antenna, the new species is related to the pallipes- species group ( Wang et al. 2007 ). Males of N. kotrbae are remarkable in having unusual setation on fore tarsus and hind tibia, strongly different from all other described species. Type material. HOLOTYPE 3 [dried and mounted on pin], La Reunion : near Ste-Rose, Foret Mourouvin, 470 m , 27.IV.2002 , leg. M. Kotrba [ ZSM ]. PARATYPES : 703 and Ƥ [in alcohol], same label [ ZSM ; 13 and 1Ƥ in author’s coll.]; 23, 3Ƥ [in alcohol], La Reunion : Le Grand Etang, Ufer, 27.IV.2002 , leg. M. Kotrba [ ZSM ]. Etymology. The species is named for the collector, Dr. Marion Kotrba. Description. Male: Head : Frons and face with black ground colour; frons wider than high, brownish pollinose. One vertical seta at upper corner of frons; one short postvertical seta positioned behind vertical seta; ocellar setae strong; two short fine postocellars; postocular setae all dark, slightly increasing downward. Eyes haired. Face under antennae slightly wider than height of postpedicel, slightly narrowing towards clypeus. Antenna black-brown, slightly longer than head height. Scape small, vase-like; pedicel larger, globular, with ring of short setulae and 1 elongate apicodorsal seta; postpedicel triangular, as long as high at base, densely haired; arista-like stylus arising from the apical concavity of the postpedicel, 2.5 times as long as main segments of antenna combined, pubescent, with short thick segment 1 and filiform segment 2. Ratio of lengths of scape to pedicel to postpedicel to stylus: 2/ 3/5/29 . Palpus and proboscis small, blackish-brown, palpus oval, with black seta and dark pubescence; proboscis with 6 pseudotracheae. FIGURE 15. Habitus of Nepalomyia kotrbae sp. nov. , after maceration. FIGURE 16. Head of Nepalomyia kotrbae sp. nov. FIGURE 17. Wing of Nepalomyia kotrbae sp. nov. FIGURE 18. Fore tarsus of Nepalomyia kotrbae sp. nov. , anterior view. FIGURE 19. Hind tibia and tarsus of Nepalomyia kotrbae sp. nov. , anterior view. FIGURE 20. Postabdomen of Nepalomyia kotrbae sp. nov. , right lateral aspect. Abbreviations: cerc, cercus; ep, epandrium; epl, epandrial lobe; hyp, hypandrium; ph, phallus; sur, surstylus; tg, tergum. FIGURE 21. Postabdomen of Nepalomyia kotrbae sp. nov. , left lateral aspect. FIGURE 22. Hypopygium of Nepalomyia kotrbae sp. nov. , left lateral aspect. FIGURE 23. Hypandrium and phallus of Nepalomyia kotrbae sp. nov. , ventral aspect. Abbreviations: hyp, hypandrium; ph, phallus. Thorax : Dark-brown, with dark bristles; mesonotum shining blue, flattened posteriorly. Five (1+4) dorsocentrals, with 5th seta slightly shifted towards lateral margin; 7−8 pairs of short acrostichals in two rows; scutellum with one pair of strong setae and one pair of lateral hairs. Upper propleuron without setae; proepisternum with 1 strong seta. Legs : Yellow with coxae and 5th tarsomeres brownish. All tarsi with simple and small pulvilli and claws. Fore and mid coxae with dark anterior and apical setae; hind coxa with 1 dark bristle at middle. Fore leg without strong setae; fore femur with 1−2 fine posteroventral subapicals, fore tarsus with anterior row of long curved cilia, 2−3 times as long as diameter of corresponding tarsomeres, 2-3 cilia on each segment, longest on basitarsus. Length ratio of fore tibia to tarsus (segments from first to fifth): 31/10/7 / 5/5/6. Mid femur simple, bearing 1 strong but short anterior subapical bristle; mid tibia simple, with 1 anterodorsal and 1 posterodorsal seta at basal third, 1 anterodorsal right below middle and 3-4 apical setae, of which one ventral longish; no ventral setae; mid tarsus simple, but basitarsus with short basoventral seta; segments 1–4 each with short apical spinules. Length ratio of mid tibia to tarsus (segments from first to fifth): 42/16/9 /8/6/6. Hind femur simple, without strong bristles; hind tibia simple, with 2 anterodorsals, posterodorsal row of 8−9 pale setae and with simple subapical dorsal bristle; hind basitarsus with 1−2 short ventrals, 1 short apicoventral seta, with very small basal tooth posteriorly. Length ratio of hind tibia to tarsus (segments from first to fifth): 45/ 10/15/8 /7/8. Wing : Greyish, veins brown. Costa simple, with simple costal setulae reaching R2+3. Ratio of part of costa between R2+3 and R4+5 to that between R4+5 and M1+2: 14/10. R2+3 and R4+5 slightly diverging towards wing apex; R4+5 and M1+2 straight and slightly diverging in distal half. Ratio of apical to basal part of M1+2: 60/35. Ratio of dm-cu to apical part of CuA1: 7/20. Anal vein fold-like, not reaching wing margin; anal lobe poorly developed; anal angle obtuse. Lower calypter small, brownish, with pale setae. Halter brownish. Abdomen : Brown, cylindrical, as long as thorax (dried specimen), with black hairs and marginal setae; tergum 6 small, glabrous, with shallow posterior emargination; sterna 5 and 6 weakly sclerotized. Segment 7 glabrous, reduced, with small sternum; in a dried specimen tergum 7 is visible as small inverted black Vshaped sclerite (dorsal view); segment 8 brown, large, ovate, setose; hypopygium brown, with yellow cercus; epandrium ovoid, longer than high, concave ventrally in middle; foramen lateral, positioned in basal half of epandrium; hypandrium not fused with epandrium, originating from 2 basolateral arms, then simple, short, parallel-sided, with rounded apex; phallus thick at base, slender, cleft at apex; epandrial lobes reduced to 3−4 simple epandrial setae, with one seta longish, and strong and long pedunculate curved seta at distoventral corner of epandrium; surstylus symmetrical, trilobate, with dorsal arm subtriangular, with dorsal seta; middle arm short, with thick apical seta; ventral arm of surstylus irregularly shaped, broad, with 2−3 small and 1 large oval plates; latter with 1 simple and 1 long flattened seta; cercus elongate-triangular, with strong apical spine and sparse dorsal setae. Measurements (mm): Body length 1.2, antenna length 0.45, wing length 1.5, wing width 0.5, hypopygium length 0.45. Female : Similar to male except lacking male secondary sexual characters, otherwise as follows. Hind tibia with 5−6 dark posterodorsals, 2 anterodorsals; fore tarsus simple; each hemitergite with 5 thick setae.