Order Chiroptera - Family Pteropodidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 313 350 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Pteropus samoensis subsp. nawaiensis Gray 1870 Synonyms: Pteropus samoensis subsp. fuscicollis Nicoll 1904 ; Pteropus samoensis subsp. ruficollis Nicoll 1908 . Discussion: samoensis species group.