Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae) Author Burks, Roger A. Department of Entomology, University of California, Riverside, CA 92521 Author Heraty, John M. Department of Entomology, University of California, Riverside, CA 92521 john.heraty@ucr.edu Author Mottern, Jason Department of Entomology, University of California, Riverside, CA 92521 & USDA Systematic Entomology Laboratory, Washington, DC 20013 - 7012 Author Dominguez, Chrysalyn Department of Entomology, University of California, Riverside, CA 92521 Author Heacox, Scott Department of Entomology, University of California, Riverside, CA 92521 text Journal of Hymenoptera Research 2017 2017-04-28 55 139 188 http://dx.doi.org/10.3897/jhr.55.11482 journal article http://dx.doi.org/10.3897/jhr.55.11482 1314-2607-55-139 0E1B0A14F8714CD89F2910B255677621 FFF5667EAD5E7A6FFFEC2B6CFFB7D800 575140 Leiosema Heraty & Burks gen. n. Figs 56 , 67 Type species. Orasema glabra Heraty, 1994: 72-73, figs 119, 125-128, 130, 132. Etymology. Combination of the Greek λεῖος ( leios ) for smooth and a contraction of Orasema , referring to the smooth and polished head and frenum; gender feminine. Discussion. Originally treated within Orasema as the glabra group by Heraty (1994 : p. 72). Heraty (1994) speculated that some of the characteristic features were shared with species in the New World coloradensis group. However, there is a greater similarity with Zuparka (Madagascar), including the entirely smooth face (Figs 57 - 58 , 62 ), scrobal channels with dorsal foveae (Fig. 57 ), labrum with 4 digits (Figs 57 , 62 ), and fore wing with bare basal area and speculum (Fig. 65 ). However, Zuparka has a laterally smooth or very weakly sculptured propodeal disc (Figs 73 , 82 ) and the pronotum and prepectus are more rigidly associated ventrally (Fig. 82 ). Australosema synempora Heraty also has the mesosomal sidelobes, axillae and frenum smooth, but the face is reticulate and the propodeum has lateral glabrous areas. Matantas also has a similar smooth face, axilla and frenum, but the face is distinctly subtriangular, the labrum has 6-8 digits and the propodeum is laterally glabrous. Each of these other genera have only 3 or 4 lateral subapical teeth on the first valvula in contrast to the 6-10 minute lateral teeth of Leiosema . Diagnosis. Separated from Orasema by the presence of male scape glands (pores) (Fig. 56 , vp), broadly impressed rugulose antecostal sulcus and the combination of smooth head, 4-digitate labrum, and smooth frenum (Figs 59 - 61 ). Distinguished from the other Old World genera by having the head smooth and subcircular in frontal view (Figs 57 - 62 ); scrobal depression with parallel channels and dorsal foveae (Fig. 57 ); dorsal occipital margin rounded; funicle 7-segmented; labrum with 4 digits; mesonotum with scattered short setae; mesoscutal lateral lobes and frenum smooth; transscutal articulation complete; propodeal disc evenly rugose; prepectus foveate and loosely articulated with pronotum ventrally (Fig. 63 ); fore wing with basal area and specular area bare (Fig. 65 ), with dense and relatively long wing disc setae; postmarginal vein 1.5-3.2 x as long as stigmal vein; petiole base truncate with prominent basal flange (Fig. 67 ); antecostal sulcus broad and reticulate; first valvula of ovipositor with 6-10 minute lateral teeth. Male scape with distinct ventral pores (glands) set in circular depressions across entire ventral surface (Fig. 56 ). Distribution. Sub-saharan Africa (Democratic Republic of the Congo, South Africa, Tanzania, Zimbabwe), and Yemen.