Four new species of Utivarachna Kishida, 1940 (Araneae: Trachelidae) from Sumatra Author Dhiya’Ulhaq, Naufal Urfi 0000-0001-6563-4643 Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany naufalurfi.du@gmail.com Author Dupérré, Nadine Centre for Taxonomy and Morphology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Museum of Nature Hamburg, Hamburg, Germany & American Museum of Natural History, New York, NY, United States of America Author Buchori, Damayanti Department of Plant Protection, Faculty of Agriculture, IPB University, Bogor, Indonesia & Centre for Transdisciplinary and Sustainability Sciences, IPB University, Bogor, Indonesia Author Scheu, Stefan Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany & Centre of Biodiversity and Sustainable Land Use, Göttingen, Germany Author Drescher, Jochen 0000-0002-5162-9779 Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany jochen.drescher@biologie.uni-goettingen.de text Zootaxa 2024 2024-03-04 5418 5 551 575 http://dx.doi.org/10.11646/zootaxa.5418.5.6 journal article 10.11646/zootaxa.5418.5.6 1175-5326 10780063 8681F446-C4A0-466F-A763-57F426B14523 Utivarachna Kishida, 1940 Type species: Utivarachna fukasawana Kishida, 1940 Diagnosis. According to Deeleman-Reinhold (2001) , the genus Utivarachna is closest to Trachelas L. Koch, 1872 but can be distinguished from it by the following characters: lateral invagination of carapace just behind the cephalic region ( Figs 1A , 3A ); pits in the margins of sternum ( Figs 1B , 3B ); triangular projection of the clypeus extending between the cheliceral bases; chitinous stripes between carapace and sternum ( Figs 1B , 3B ); lengthened leg I with multiple rows of ventral cuspules in males ( Figs 1C , 7C ) (versus not lengthened and with one row of cuspules or cuspules absent); embolus forming transverse apical coil ( Figs 2 , 4 ); epigynum with large posterior atrium ( Figs 6 , 9D–E ); copulatory openings located posteriorly (versus more anteriorly-located, Figs 6 9D–E ). Composition. Utivarachna contains 31 species that are currently divided into four species groups. For their diagnosis, see Deeleman-Reinhold (2001) and Liu et al . (2020) : dusun -group: U. dusun Deeleman-Reinhold, 2001 ; U. itiokai Yamasaki, 2023 ; U. rubra Deeleman-Reinhold, 2001 and U. yumaoi Lin & Li, 2023 . fukasawana -group: U. chamaeleon Deeleman-Reinhold, 2001 ; U. galyanae Dankittipakul, Tavano & Singtripop, 2011 ; U. fukasawana Kishida, 1940 and U. ichneumon Deeleman-Reinhold, 2001 . kinabaluensis -group: U. accentuata ( Simon, 1896 ) ; U. angsoduo sp. nov. ; U. arcuata Zhao & Peng, 2014 ; U. bucculenta Deeleman-Reinhold, 2001 ; U. convolutiva Dankittipakul, Tavano & Singtripop, 2011 ; U. fabaria Zhao & Peng, 2014 ; U. fanjing Li, Zhang & Yu, 2022 ; U. fronto ( Simon, 1906 ) ; U. gongshanensis Zhao & Peng, 2014 ; U. kinabaluensis Deeleman-Reinhold, 2001 ; U. lata Jin, Yin & Zhang, 2015 ; U. linyejiei Chu & Li, 2023 ; U. rama Chami-Kranon & Likhitrakarn, 2007 ; U. subfabaria Liu, Xu & Haddad, 2020 ; U. taiwanica ( Hayashi & Yoshida, 1993 ) ; U. tamdao Chu & Li, 2023 ; U. tangi Liu, Xu & Haddad, 2020 ; U. trisula sp. nov. and U. zhengguoi Chu & Li, 2023 . phyllicola -group: U. balonku sp. nov. ; U. gui (Zhu, Song & Kim, 1998); U. phyllicola Deeleman-Reinhold, 2001 and U. rimba sp. nov. Distribution. Tropical Asia: India , Sri Lanka , China , Taiwan , Myanmar , Vietnam , Thailand , Peninsular Malaysia , Sumatra, and Borneo.