Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini) Author Ocampo, Federico C. text The Coleopterists Bulletin 2004 2004-06-30 58 2 295 305 http://dx.doi.org/10.1649/685 journal article 10.1649/685 1938-4394 10104682 Glyphoderus monticola Burmeister, 1861:67 . Diagnosis. Males. This species is separated from other Glyphoderus species by the shape and placement of the pronotal horns and tubercles ( Fig. 4 ). Glyphoderus monticola has the pronotum with two horns in middle and two transversally elongated tubercles (sometimes developed as horns) on the basal half; horns and tubercles on margin of medial depression; anteromedial horn with ‘‘arms’’ elongated, with two anteromedial teeth; distance between teeth as long as tooth base. No morphological characters were found to diagnose females of this species, females can be identified only by association with males. Distribution ( Fig. 6 ). ARGENTINA : Catamarca : El Recreo (3). La Rioja : La Rioja (3); La Rioja ( 20 km N) (4); Mascasin (33); no more data (2). Temporal Distribution. January (31), February (4), March (3), November (2), December (1).