Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)
Author
Lazányi, Eszter
Author
Vagalinski, Boyan
text
Zootaxa
2013
3741
1
55
100
journal article
10.11646/zootaxa.3741.1.2
6a6ba78e-a9c5-4d11-997a-3d06e5e31520
1175-5326
223773
BF5EA9B8-C6F4-448A-BEF9-1976AB4EC308
Key to males of subgenus
Megaphyllum
sensu
stricto
on the basis of gonopod characters and body colour patterns
1 Anterior process (
ap
) well discernible.................................................................... 2
- Anterior process (
ap
) reduced to a low ridge or fully absent................................................... 5
2 Posterior process (
pp
) shorter than solenomere............................................................. 3
- Posterior process (
pp
) longer or subequal to solenomere...................................................... 4
3 Anterior process (
ap
) perpendicular to the main opisthomere body, with toothed apex (
Fig. 5d
)..............
M. silvaticum
- Anterior (
ap
) process parallel to the main opisthomere body, with normal blunt apex (
Figs 4d, 4f
)...........
M. projectum
4 Posterior solenomere process (
psp
) large, massive, standing high above the anterior solenomere process (
asp
) (
Fig. 2d
). Dark grey animals with very wide reddish longitudinal band.........................................
M. erythronotum
- Posterior solenomere process (
psp
) subequal to the anterior solenomere process (
asp
) (
Fig. 3d
). Reddish-brown animals with two bright longitudinal dorsal bands............................................................
M. metsovoni
5 Apical anterior hump (
ah
) well developed into a more or less blunt protuberance in front of the anterior solenomere process (
asp
)............................................................................................... 6
- Apical anterior hump (
ah
) not detectable................................................................ 11
6 Promere with a unique rod-like process below its apex (
Fig. 14c
). Dark grey animals with two bright dorsal longitudinal bands; uniformly dark forms also present........................................................
M. transsylvanicum
- Promere with normal apex............................................................................. 7
7 Posterior process (
pp
) joining the main opisthomere body at a relatively wide angle of ca. 50˚ (
Fig. 17b
). Promere strongly tapering (
Fig. 17c
).......................................................................
M. spathulatum
- Posterior process (
pp
) joining the main opisthomere body at a more acute angle of not more than 20˚–30˚. Promere more grad- ually narrowing..................................................................................... 8
8 Posterior process (
pp
) shorter than solenomere (
Fig. 16d
). Uniformly dark brown animals without longitudinal lines...............................................................................................
M. rhodopinum
- Posterior process (
pp
) subequal to solenomere............................................................. 9
9 Anterior solenomere process (
asp
) more than two times longer than the posterior solenomere process (
psp
) (
Fig. 19d
). Plain brownish animals............................................................................
M. tauricum
- Anterior solenomere process (
asp
) just slightly exceeding the posterior solenomere process (
psp
).................... 10
10 Posterior solenomere process (
psp
) enlarged, thicker than the anterior solenomere process (
asp
) (
Fig. 17b
). Promere posteri- orly with a large lateral lobe (
l
on
Figs 17a, 17c
). Brown animals with two bright longitudinal bands......
M. sjaelandicum
- Posterior solenomere process (
psp
) normal, maybe only slightly ticker than the anterior solenomere process (
asp
) (
Fig. 17b
). Promere without a prominent lateral lobe. Dark grey animals with one bright dorsal longitudinal line......
M. unilineatum
11 Anterior solenomere process (
asp
) strongly elongated, surpassing by far the posterior solenomere process (
psp
)......... 12
- Anterior solenomere process (
asp
) shorter or subequal to the posterior solenomere process (
psp
)..................... 13
12 Posterior process (
pp
) subequal to solenomere, apically serrated, as the anterior solenomere process (
asp
) (
Fig. 14d
*). Dark grey animals with two yellowish dorsal longitudinal bands............................................
M. kievense
- Posterior process (
pp
) much shorter than solenomere, apically smooth like the anterior solenomere process (
asp
) (
Fig. 13b
). Males uniformly dark grey with reddish legs; females with two bright longitudinal dorsal bands..........
M. rosenauense
13 Posterior solenomere process (
psp
) much longer and wider than the anterior solenomere process (
asp
)................ 14
- Posterior solenomere process (
psp
) subequal to the anterior solenomere process (
asp
).............................. 17
14 Promere apically with a large toothed or serrated lobe (
sl
on
Fig. 9a
) and a shelf-like protrusion from the distal part of its ante- rior surface (
Fig. 9c
).........................................................................
M. dentatum
- Promere without special characters..................................................................... 15
15 Posterior solenomere process (
psp
) swan-neck shaped; posterior process (
pp
) higher than solenomere (
Fig. 8d
)....................................................................................................
M. cygniforme
- Posterior solenomere process (
psp
) differently shaped; posterior process (
pp
) subequal to solenomere................. 16
16 Posterior solenomere process (
psp
) wavy (
Fig. 11d
). Uniformly dark brown animals...................
M. montivagum
- Posterior solenomere process (
psp
) smooth (
Fig. 10d
).............................................
M. glossulifer
17 Posterior process (
pp
) surpassing solenomere............................................................. 18
- Posterior process (
pp
) shorter or subequal to solenomere..................................................... 19
18 Posterior process (
pp
) wide and rounded, fully concealing solenomere from posterior view (
Fig. 12a
); posterior solenomere process (
psp
) smooth (
Fig. 12d
). Dark grey animals with yellow to reddish dorsal side, usually with a thin blackish mid-dorsal line.....................................................................................
M. bosniense
- Posterior process (
pp
) slender, strongly tapering; posterior solenomere process (
psp
) wavy (
Fig. 7b
). Dark brown or grey animals with thin mid-dorsal black line...........................................................
M. carniolense
19 Posterior process (
pp
) wide and rounded; anterior and posterior solenomere processes (
asp
and
psp
) slender, finger-like (
Fig. 15d
). Very large dark grey animals with narrow, yellow to reddish longitudinal dorsal band..................
M. hercules
- Posterior process (
pp
) narrow, apically pointed; anterior and posterior solenomere processes (
asp
and
psp
) rather short, weakly protruding (Fig. 6d)........................................................................
M. austriacum