A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia Author Zonstein, Sergei Author Kunt, Kadir B. Author Yağmur, Ersen A. text European Journal of Taxonomy 2018 2018-01-30 399 1 93 journal article 30873 10.5852/ejt.2018.399 b389e1f3-82be-4258-9b09-b7ef9d839f99 1163422 E836E138-D6E2-4F62-B4B3-CE2E073F2B24 Raveniola birecikensis sp. nov. urn:lsid:zoobank.org:act:1DB28F4C-3186-4E0E-9769-C533F62AB5D4 Figs 50 , 72–73 , 106 , 133–134 , 153–154 , 180–181 , 221 Diagnosis The male of this species differs from those of other members of the same group by having a welldeveloped keel in the flattened proximal part of the embolus ( Figs 180–181 ; cf. Figs 183–188 ). Raveniola biresecensis sp. nov. can be distinguished from R. nana sp. nov. (male of the latter species is unknown) by its noticeably larger body size, more intensely brownish coloration, and relatively smaller AMEs and PMEs ( Fig. 106 ; cf. 122). Etymology The specific epithet is a toponym that refers to the locality where the paratypes were collected: Birecik District, Şanlıurfa Province of Turkey. Material examined Holotype TURKEY : ♂ , Şanlıurfa Province , Halfeti District , Şanlıurfa, N of Halfeti , 37°16′ N , 37°51′ E , 400– 600 m , 7 Mar. 2006 , A.E. Yağmur leg. ( AUZM ). Paratypes TURKEY: 2 ♂♂, Şanlıurfa Province, Birecik District, Yukarı-Habip, 11 km of NNE Birecik, 37°04.2′ N, 38°00.5′ E, 550–650 m, 1 Apr. 2006, A.E. Yağmur leg. (AUZM, TAU). Aditional material (5 possibly dwarf ♂♂) TURKEY: 3 ♂♂, Şanlıurfa Province, Halfeti District, 37°16′ N, 37°51′ E, 400–600 m, 14 Feb. 2011, E.A. Yağmur leg. (AUZM and TAU); 2 ♂♂, surroundings of Birecik, 37°04′ N, 38°00′ E, 550–650 m, 4 Feb. 2011, E.A. Yağmur leg. (AUZM, TAU). Description Male (holotype) HABITUS. See Fig. 50. MEASUREMENTS. TBL 14.77, CL 4.66, CW 4.08, LL 0.28, LW 0.66, SL 2.26, SW 2.02. COLOUR. Carapace, palps and legs dorsally light yellowish brown, with darkened cephalic area, thoracic grooves and leg I; eye tubercle dark brown; chelicerae medium cherry-red; sternum, labium, maxillae, palps and legs ventrally light yellowish brown; abdomen dorsally light yellowish grey, with indistinct darker dorsal pattern, ventral abdominal surface and spinnerets pale yellowish grey. PROSOMA. Carapace and chelicerae as shown in Fig. 72 . Clypeus and eye group as in Fig. 106 . Eye diameters and interdistances: AME 0.12(0.16), ALE 0.20, PLE 0.14, PME 0.07, AME–AME 0.19(0.16), ALE–AME 0.10(0.08), ALE–PLE 0.06, PLE–PME 0.05, PME–PME 0.44. Each cheliceral furrow with 9 promarginal teeth and 9–10 mesobasal denticles. Maxillae with 7 relatively small cuspules each. LEGS. Tibia and metatarsus I as shown in Fig. 133 . Scopula: distal on metatarsi I–II, entire on tarsi I–II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–11 each on tibiae, 15–17 on metatarsi, 10 on tarsi I and II, 10–12 on tarsi III and IV, 7 on cymbium. Paired claws with 8–10 teeth on each margin. LEG MEASUREMENTS. Femur Patella Tibia Metatarsus Tarsus Total SPINATION. Femora I–IV with one basodorsal spine and 3 distal bristles; patellae I and II aspinose. Palp: femur d1–1–1–1–1, pd1; patella p1; tibia d1–1, p1–1–1, r1, v1–1; cymbium d4(5). Leg I: femur pd0–0(1)–1; tibia rv0–1–m–m, v1–1(0)–0l; metatarsus v0–0–2. Leg II: femur pd0–0–1; tibia p0–0–1, v2–2–3; metatarsus p1, v2–2–3. Leg III: femur pd0–1–1, rd0–1–1; patella p1, r1; tibia d1–0–0, p1– 1(0)–1, r1–1–1, v2–2–3; metatarsus d0–1–0, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur pd0–1–1, rd0–0– 1; patella p1, r1; tibia d1–0–0, p1–1–1, r1–1–1, v2–2–3; metatarsus d0–1–1–0, p1–1–1–1, r1–1–1–1, v2–2–2–3.

Palp	2.40	1.35	1.88	–	0.84	6.47
Leg I	3.98	2.29	3.04	3.07	1.63	14.01
Leg II	3.61	2.17	2.67	2.68	1.64	12.77
Leg III	2.92	1.76	1.89	2.76	1.58	10.91
Leg IV	3.66	2.16	3.27	4.30	1.74	15.13

PALP. Tibia, cymbium and bulb as shown in Fig. 153 . Wide and flattened basal part of embolus with subapical keel extending into twisted and flagelliform apical part ( Figs 180–181 ). SPINNERETS. PMS: length 0.49; diameter 0.13. PLS: maximal diameter 0.41; length of basal, medial and apical segments 0.67, 0.42, 0.47; total length 1.56; apical segment shortly digitiform. Female Unknown. Variation Carapace length in males of the type series varies from 3.93 to 4.66. Colouration, eye arrangement, shapes of the palpal tibia, cymbium, palpal organ, tibia and metatarsus I in the paratypes do not differ significantly from those of the holotype ( Figs 73 , 134 , 154 ). The appearance and the corresponding structures of the pale-coloured and possibly dwarf conspecific males (CL 2.16–3.41), possessing larger AMEs and an invariably shorter basal portion of the embolus, are shown in Figs 47 , 74 , 107 , 135 , 155 and 182 , respectively. Ecology The type series, which includes only males, was collected with pitfall traps in open steppe biotopes sparsely covered with low forest and shrubs ( Fig. 230 ). Distribution The species is known from south-eastern Turkey (Şanlıurfa Province) and probably also occurs in the adjoining territory of Syria. See Fig. 221 .