The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea)
Author
Shear, William A.
0000-0002-5887-7003
Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA. wshear @ hsc. edu; https: // orcid. org / 0000 - 0002 - 5887 - 7003
wshear@hsc.edu
Author
Marek, Paul E.
Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.
text
Zootaxa
2022
2022-11-11
5205
6
501
531
journal article
187956
10.11646/zootaxa.5205.6.1
4b6eb79c-018f-470b-ac46-6646270ec7ee
1175-5326
7318605
9D1DEC25-5FA2-4D64-807E-F103C3FCB5CA
Nototrisaria ornata
Shear & Marek
,
n. sp.
Figs 1–6
,
49–51
FIGURE 1.
Nototrisaria ornata
,
n. gen., n. sp
.
, live female in habitat. About 5.0 mm long. Photo by Bill Leonard.
FIGURES 2–6.
Nototrisaria ornata
,
n. gen., n. sp
.
2.
Anterior end of male, ventral view.
3.
Rings 2–4 of male, dorsal view.
4.
Posterior end of male, oblique view.
5.
Right gonopod, lateral view.
6.
Legpair 9 of male, anterior view. Abbreviations:
aac
, anterior angiocoxite;
col
, collum;
cc
, colpodoxite;
c1,
crest 1;
c6
, crest 6;
cxs
, coxosternite;
cx10
, coxa of leg 10;
L1
, leg 1;
L2
, leg 2;
L3
, leg 3;
m
, mandible;
numerals
, ring numbers;
pac
, posterior angiocoxite;
t9
, telopodite of leg 9.
Types:
Male
holotype
and male
paratype
from
9.6 miles
up
Elochoman
Valley
Road
from Rt. 4,
Wahakiakum Co.
,
Washington
,
46.289633°N
, -
123.26485°W
, 450’ asl, collected
28 March 2004
by
C. Richart
,
W. Leonard
;
male
paratype
from
11.7 mi
up
Elochoman
Valley
Road
from
Rt.
4, 450’ asl,
46.316783°N
, -
123.2617°W
,
28 March 2004
, collected by
C. Richart
;
male and female
paratypes
from
Trap Creek
,
1.1 mi
S of SR6,
Pacific Co.
Washington
,
46.5468°N
, -
123.6161°W
, 180’ asl, collected
11 January 2003
and
3 May 2003
by
W. Leonard.
Parts of a male
paratype
are mounted on SEM stub WS3-8.
Types
deposited in the CAS.
Etymology
: The species name, a Latin adjective, refers to the ornate dorsum.
Diagnosis
: As for the genus; see above.
Description
:
Male
paratype
from Elochoman River Road.
Length,
5.5 mm
, width
0.6 mm
(includes paranota). Thirty postcephalic rings including telson.
Head (
Fig. 2
) densely setose, with sculpture of tiny, acute tubercles; labral corners rounded, without projecting hook. Ommatidia absent. Antennae short, robust. Mandibular stipes (
Fig. 2
, m) with sawtooth distal edge, acute apical projection. Color of live specimen white to pale yellow (
Fig. 1
).
Collum (
Fig. 2
, col) broadly expanded, with 10 subequal crests limited to posterior two-thirds; anterior third with pebbled sculpture. Anterior rings (
Fig. 3
) with C2 and C6 exaggerated, the latter expanded into prominent paranota with deep posterior notch; C1 reduced to a few knobs, absent on more posterior rings, C2 also progressively lower on midbody ring, exaggerated again on most posterior 6–8 rings (
Fig. 4
). Metazonital setae not observed, probably absent. Limbus serrate.
Telson without crests, lobes deeply incised (
Fig. 4
).
First legpair crassate (
Fig. 2
, L
1
), larger than legpairs 2 and 3 (
Fig. 2, L2, L3
), postfemora and tibia set with long, stiff setae, tarsus with comb. Legpair 2 relatively small, similar to postgonopodal legs. Legpair 3 with prominent, broad, slightly sigmoidal coxal flasks bearing curled setae on anterior surface (
Fig. 2
, cf); prefemora expanded, flattened. Legpairs 4–6 strongly crassate. Legpair 7 similar to pairs 4–6, but with large, flattened coxal plates covering bases of gonopods.
Gonopods in mesal view (
Figs 5
,
50
) with anterior angiocoxites erect, narrow, hooked and expanded at tip; posterior edge with continuous fimbriate lamina, lamina ends in long distal projection; tuft of strong, thin cuticular projections subapically (aac,
Figs 5
,
49, 50
). Posterior angiocoxites as broad, rounded plate (pac,
Figs 5
,
50
). Flagellocoxites absent. Colpocoxite (
cc,
Figs 5
,
49, 50
) developed as compact mass of finely divided, sinuous fimbriae with small posterior portion unmodified, sclerotized.
Ninth legpair (
Figs 6
,
51
) strongly reduced, sternum (s9), coxae (cx) and telopodites (t9) fused; coxal part with blunt triangular process (cp), telopodite part with long, finger-like basal process bearing acute setae.
Tenth leg with enlarged, glandular coxae (
Fig 6
, cx10).
Female
paratype
similar to male in nonsexual characters, but exaggerated crests significantly lower. A living female specimen is depicted in
Fig. 1
.
Distribution
: As for the genus. The Grays Harbor County record (
WASHINGTON
: Grays Harbor Co.: Canyon River,
47.303517°N
, -
121.508167°W
, 1500’ asl,
12 December 2004
, W. P. Leonard) is based on a female that strongly resembles the females from other localities.