Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 12) Phortica ( Ashima ) foliiseta Duda, 1923 Phortica foliiseta Duda, 1923: 35 ; Chen et al ., 2005b: 420 . Amiota ( Phortica ) foliiseta : Tsacas & Okada, 1983: 230 . Phortica ( Ashima ) foliiseta : Chen & Máca, 2012: 507 . Diagnosis. Dorsomedial, tentorial apodeme 3/4 as long as basal, parallel portion of dorsolateral, tentorial apodeme; longest, dorsal branch of arista as long as longest seta on pedicel; each lateral lobe of aedeagal sheath with apically more or less pointed, sclerotized, anterior and posterior postgonites (“ Fig. 4 ” in Tsacas & Okada 1983 ). Supplementary description. Supracervical setae approximately 12. Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0). Arista with arb = 1–2/0; apical, leaf-like expansion narrower than long, apically tapering and pointed (ch.7-1). Cibarial, medial sensilla approximately 7 per side; posterior sensilla approximately 6 per side. Fore and mid tibiae entirely yellow (ch.17,18-0). All tarsi entirely yellow. Abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0), without dense setulae on medial surface (ch.26-0). Pregonite gently curved ventrad distally. Posterior postgonites on lateral lobes of aedeagal sheath medially separated from each other (ch.66-0). The antisymmetry is observed in the postgonites: in A-type, the right, posterior postgonite is slightly curved but the left one nearly straight, and the right, anterior postgonite is thicker than the left one (“ Fig. 4 ” in Tsacas & Okada 1983 ); but in B-type, vice versa . Specimens examined. Taiwan : 2♂ (A-type), Shanlinxi , 16.x.1992 , M.J. Toda leg. ( SEHU ) ; 3♂ (B-type), Shan- linxi, 15–16.x.1992 , M.J. Toda leg. ( SEHU ) ; 1♂ (A-type), Wulai , 8.x.1992 , M.J. Toda leg. ( SEHU ) . China : 1♂ ( Btype ), Dinghushan , Guangdong , 4–10.iii.1987 , T .- X. Peng leg. ( SEHU ) . Distribution. Taiwan , China ( Guangdong ), Thailand . Remarks. This species and the subsequent three species form a cluster (BP = 88%) with three synapomorphies ( Fig. 1 ): abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0), without dense setulae on medial surface (ch.26-0). However, this species can be distinguished from them by the diagnostic characters.