<document id="B7B4370ABB50F6890A41381AC3A6B6D6" ID-CLB-Dataset="9024" ID-DOI="10.11646/zootaxa.4861.4.4" ID-GBIF-Dataset="db44bc90-2960-490c-8ddb-f16633c03929" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416747" ID-ZooBank="2297FB97-3C02-42B0-B811-019646E33C0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609801195912" checkinUser="plazi" docAuthor="Smales, Lesley" docDate="2020" docId="0B7B7959C463F810FF60FB62AF0CF9B5" docLanguage="en" docName="zootaxa.4861.4.4.pdf" docOrigin="Zootaxa 4861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helgenema Smales 2020, new genus" docType="treatment" docUuid="A5E1C6BE-9D63-47AB-AF48-E2FA4E982F3E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="553" masterDocId="F7420121C46AF819FFF7FF90AD0EFF98" masterDocTitle="Gastrointestinal nematodes of Paramelomys levipes and P. mollis (Rodentia Muridae) from Papua, Indonesia and Papua New Guinea with the descriptions of three new genera and nine new species (Nematoda)" masterLastPageNumber="572" masterPageNumber="544" pageNumber="553" updateTime="1698852240458" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A9946C8CC6028051A5BA8FE37472784F">Gastrointestinal nematodes of Paramelomys levipes and P. mollis (Rodentia Muridae) from Papua, Indonesia and Papua New Guinea with the descriptions of three new genera and nine new species (Nematoda)</mods:title>
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<mods:date id="DAEACFC7BA95EBD30E86D01E4A34CF97">2020</mods:date>
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<taxonomicName id="44D2B3CCC463F810FF60FB62AC2CFA95" authority="Smales, 2020" authorityName="Smales" authorityYear="2020" box="[151,290,1266,1293]" class="Mammalia" family="Muridae" genus="Helgenema" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="553" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B1A6145DC463F810FF60FB62AC2CFA95" bold="true" box="[151,290,1266,1293]" italics="true" pageId="9" pageNumber="553">Helgenema</emphasis>
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<emphasis id="B1A6145DC463F810FF60FAAAAC01FACC" bold="true" box="[151,271,1338,1364]" pageId="9" pageNumber="553">Definition</emphasis>
. 
<taxonomicName id="44D2B3CCC463F810FEE8FAABACD3FACD" box="[287,477,1339,1365]" class="Chromadorea" family="Heligmonellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strongylida" pageId="9" pageNumber="553" phylum="Nematoda" rank="family">Heligmonellidae</taxonomicName>
, 
<taxonomicName id="44D2B3CCC463F810FE1DFAABAFB0FACD" box="[490,702,1339,1365]" class="Chromadorea" family="Heligmonellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strongylida" pageId="9" pageNumber="553" phylum="Nematoda" rank="subFamily" subFamily="Nippostrongylinae">Nippostrongylinae</taxonomicName>
. Synlophe well developed with 11–15 pointed longitudinal continuous ridges. Cuticle with dilatation on left side supported anteriorly by large left dorso-lateral ridge. Axis of orientation sub frontal. Bursal pattern 2–3. Dorsal ray divided within distal half. Spicule to body length ratio 7.2%. Parasites of uromyin murids.
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<typeStatus id="5C6976EDC463F810FF30FA5BADF1FA7D" box="[199,255,1483,1509]" pageId="9" pageNumber="553">Type</typeStatus>
species 
<taxonomicName id="44D2B3CCC463F810FEA9FA5BAF3BFA7D" authorityName="Smales" authorityYear="2020" box="[350,565,1483,1509]" class="Mammalia" family="Muridae" genus="Helgenema" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="keablei">
<emphasis id="B1A6145DC463F810FEA9FA5BAF3BFA7D" box="[350,565,1483,1509]" italics="true" pageId="9" pageNumber="553">Helgenema keablei</emphasis>
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<emphasis id="B1A6145DC463F810FF30FA7FAC40F991" bold="true" box="[199,334,1519,1545]" pageId="9" pageNumber="553">Etymology.</emphasis>
The genus name is in recognition of the work of Dr Kristofer Helgen who has studied extensively the murid fauna of New 
<collectingCountry id="FBC588DFC463F810FE5DF983ACF1F9B5" box="[426,511,1555,1581]" name="Guinea-Bissau" pageId="9" pageNumber="553">Guinea</collectingCountry>
.
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