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<mods:title id="A250D707E16DDECD4367A66F0958320C">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<taxonomicName id="1381E691FFC9FFEBFF2D53321D9FF8B9" ID-CoL="5X9FL" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,901,1833,1858]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="hespenheidei" subGenus="Canthidium">
<emphasis id="E6F54100FFC9FFEBFF2D53321D9FF8B9" bold="true" box="[151,901,1833,1858]" pageId="8" pageNumber="459">
<emphasis id="E6F54100FFC9FFEBFF2D53321F38F8B9" bold="true" box="[151,290,1833,1858]" italics="true" pageId="8" pageNumber="459">Canthidium</emphasis>
(
<emphasis id="E6F54100FFC9FFEBFE8853321FA7F8B9" bold="true" box="[306,445,1833,1858]" italics="true" pageId="8" pageNumber="459">Canthidium</emphasis>
) 
<emphasis id="E6F54100FFC9FFEBFE7653321C79F8B9" bold="true" box="[460,611,1833,1858]" italics="true" pageId="8" pageNumber="459">hespenheidei</emphasis>
<bibRefCitation id="B010E0E3FFC9FFEBFDD053321D9FF8B9" box="[618,901,1833,1858]" pageId="8" pageNumber="459" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
</emphasis>
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<emphasis id="E6F54100FFC9FFEBFF7D53561FFDF89E" box="[199,487,1869,1893]" italics="true" pageId="8" pageNumber="459">Canthidium hespenheidei</emphasis>
<treatmentCitation id="5520BB03FFC9FFEBFE4A53561D2DF89D" author="" box="[496,823,1869,1894]" page="92" pageId="8" pageNumber="459">
<bibRefCitation id="B010E0E3FFC9FFEBFE4A53561D14F89D" box="[496,782,1869,1894]" pageId="8" pageNumber="459" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
: 92
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</taxonomicName>
[here transferred to the subgenus 
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<emphasis id="E6F54100FFC9FFEBFB7953561B5DF89E" box="[1219,1351,1869,1893]" italics="true" pageId="8" pageNumber="459">Canthidium</emphasis>
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]. 
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locality:
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<collectingCountry id="AC96DD82FFC9FFEBFF4453691F41F872" box="[254,347,1906,1929]" name="Panama" pageId="8" pageNumber="459">Panama</collectingCountry>
</collectingRegion>
: 
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<collectingCountry id="AC96DD82FFC9FFEBFEDC53691FD9F871" box="[358,451,1906,1930]" name="Panama" pageId="8" pageNumber="459">Panamá</collectingCountry>
</collectingRegion>
: Cerro Campana, 
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, 
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, 
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. 
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<typeStatus id="0B3A23B0FFC9FFEBFC05536A1DE0F871" box="[959,1018,1905,1930]" pageId="8" pageNumber="459">Type</typeStatus>
material:
</emphasis>
<typeStatus id="0B3A23B0FFC9FFEBFBCC536A1AC4F871" box="[1142,1246,1905,1930]" pageId="8" pageNumber="459" type="holotype">Holotype</typeStatus>
male (CMNCex Howden collection); see full list of 
<typeStatus id="0B3A23B0FFC9FFEBFDE2538D1CD9F855" box="[600,707,1942,1966]" pageId="8" pageNumber="459" type="paratype">paratypes</typeStatus>
in 
<treatmentCitation id="5520BB03FFC9FFEBFD57538E1A59F855" author="" box="[749,1091,1941,1966]" page="93" pageId="8" pageNumber="459">
<bibRefCitation id="B010E0E3FFC9FFEBFD57538E1A59F855" box="[749,1091,1941,1966]" pageId="8" pageNumber="459" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young (1981: 93)</bibRefCitation>
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. 
<emphasis id="E6F54100FFC9FFEBFBEB538E1AF6F855" bold="true" box="[1105,1260,1941,1966]" pageId="8" pageNumber="459">Distribution:</emphasis>
<collectingRegion id="164553F0FFC9FFEBFB4C538E1B54F855" box="[1270,1358,1941,1966]" country="Mexico" name="Mexico" pageId="8" pageNumber="459">
<collectingCountry id="AC96DD82FFC9FFEBFB4C538E1B54F855" box="[1270,1358,1941,1966]" name="Mexico" pageId="8" pageNumber="459">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="AC96DD82FFC9FFEBFAE6538D1ED7F829" name="Costa Rica" pageId="8" pageNumber="459">Costa Rica</collectingCountry>
, 
<collectingRegion id="164553F0FFC9FFEBFF6253A11F2EF82A" box="[216,308,1978,2001]" country="Panama" name="Panama" pageId="8" pageNumber="459">
<collectingCountry id="AC96DD82FFC9FFEBFF6253A11F2EF82A" box="[216,308,1978,2001]" name="Panama" pageId="8" pageNumber="459">Panama</collectingCountry>
</collectingRegion>
, and 
<collectingCountry id="AC96DD82FFC9FFEBFED453A21FD1F829" box="[366,459,1977,2002]" name="Ecuador" pageId="8" pageNumber="459">Ecuador</collectingCountry>
(Kohlmann &amp; Solís 2006a).
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