Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains Author Wang, XIN-PING Author Griswold, CHARLES E. Author Miller, JEREMY A. text Zootaxa 2010 2593 1 127 http://www.mapress.com/zootaxa/2010/f/zt02593p127.pdf journal article zt02593p127 Draconarius pseudocapitulatus Wang 2003 (Figs 407-427, 547) Draconarius pseudocapitulatus Wang 2003: 545, figs 53A-B ( female holotype and paratypes from Gongshan County , Yunnan , China , in HNU and CAS , examined) . Additional material examined: CHINA : Yunnan : Gongshan County : 1♀ , Cikai Township, Dabadi troops place along river banks, 3030-3045 m , N27°47' , E98°30' , October 2-3, 2002 , night, October 8-10, 2002 , H.M. Yan ( CAS , CASENT9025828 ) ; 1♀ , Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi both sides , N27°47' , E98°32' , 3100 m , September 30, 2002 , H.M. Yan ( HNU , CASENT9025840 ) ; 2♀ , Bing Zhong Luo Township, Chu Kuai , N27.97928° , E098.47389° , 3725 m , August 19, 2006 , P. Hu ( CAS , CASENT9025578 ) . Fugong County : 1♀ , Pihe Township, Yueliangtian Village , N26.56784° , E098.90884° , 1520 m , August 20-24, 2005 , G. Tang , ( HNU , CASENT9025612 ) ; 8♀ , Lishadi Township, Yakou of Shibali , N27.21234° , E098.69601° , 3615 m , August 5-7, 2005 , G. Tang ( CAS , CASENT9025581 ) ; 1♀ , Lishadi, 1 km before Shibali Yaku (pass #31), 900m N of the road , N27.21447° , E98.70064° , 3585 m , August 12, 2005 , P. Paquin ( HNU , CASENT9022555 ) ; 1♀ , Lishadi, 500 m before Shibali Yaku (pass #31) , N27.21354° , E98.70021° , 3585 m , stable scree slope on soil, August 7, 2005 , P. Paquin ( HNU , CASENT9022553 ) ; 1♀ , Lishadi, Shibali , N27.16650° , E98.77936° , 2537 m , forest, webs on tree trunks at night, August 5, 2005 , P. Paquin , D. Kavanaugh ( CAS , CASENT9022568 ) ; 1♀ , Lishadi, 500 m W of Shibali , N27.16650° , E98.77936° , 2537 m , deciduous forest litter, August 4, 2005 , P. Paquin ( CAS , CASENT9022579 ) ; 1 ♀, Lishadi, 8.5 km W of Shibali , N27.18365° , E98.72002° , 3030 m , along river, on slope 3-10 m away from the river, under rocks in stable portion, August 8, 2005 , P. Paquin , D. Kavanaugh ( HNU , CASENT9022541 ) ; 1 ♀, Lishadi, 500 m W of Shibali , N27.16650° , E98.77936° , 2537 m , deciduous forest litter, August 4, 2005 , P. Paquin ( CAS , CASENT9022578 ) ; 1♀ , Lishadi, 10.5 km W of Shibali, 100 m N road , N27.20192° , E98.71321° , 3250 m , dead conifer logs in conifer forest with rhododendron patch, August 17, 2005 , P. Paquin ( HNU , CASENT9022566 ) ; 1♀ , Lishadi, 500 m before Shibali Yaku (pass #31) , N27.21354° , E98.70021° , 3585 m , stable scree slope on soil, August 7, 2005 , P. Paquin ( CAS , CASENT9022552 ) ; 2 ♀, Gaoligongshan, Shibali Forest Station , in moist stream gully, N27.16894° , E98.77205° , 2650 m , May 9, 2004 , C. Griswold & H.M. Yan ( CAS , CASENT9019907 ) ; 4♀ , Gaoligongshan, 6.18 km 280° W of Shibali , turning rocks in open meadow along stream, N27.18413° , E98.72024° , 3100 m , May 7, 2004 , C. Griswold & D. Kavanaugh ( HNU , CASENT9020708 ) ; 4♀ , Gaoligongshan, 6.18 km W of Shibali , turning rocks in open meadow along stream, N27.18413° , E98.72024° , 3100 m , May 7, 2004 , C. Griswold & D. Kavanaugh ( CAS , CASENT9020707 ) ; 1♀ , Gaoligongshan, 7.61 km directly WNN of Shibali , Rhododendron thicket, turning rocks amid bamboo, N27.20662° , E98.71773° , 3292m , May 6, 2004 , C. Griswold ( HNU , CASENT9020787 ) ; 1♀ , Gaoligongshan, 0.5 km radius of Shibali Forest Station , N27.16519° , E98.77891° , 2525 m , searching at night in forest , May 2-5, 2004 , C. Griswold & D. Kavanaugh ( CAS , CASENT9020027 ) ; 1♀ , Gaoligongshan, Shibali Forest Station , in moist stream gully, N27.16894° , E98.77205° , 2650 m , May 9, 2004 , C. Griswold & H.M. Yan ( HNU , CASENT9019908 ) ; 1♀ , Gaoligongshan, 7.61 km directly WNN of Shibali , Rhododendron thicket, turning rocks amid bamboo, N27.20662° , E98.71773° , 3292 m , May 6, 2004 , C. Griswold ( CAS , CASENT9020786 ) ; 1♀ , LiShadi subdistrict, Gaoligongshan, 12 km from Shibali toward Yaping Yakou , N27.19980° , E98.71375° , 3200 m , May 5, 2004 , H.M. Yan ( HNU , CASENT9020616 ) ; 1♂ 1♀ , Gaoligongshan, 6.18 km W of Shibali , turning rocks in open meadow along stream, N27.18413° , E98.72024° , 3100 m , May 7, 2004 , C. Griswold & D. Kavanaugh ( CAS , CASENT9020658 ) ; 1♂ , Pihe Township, Yueliangtian Village , N26.56784° , E 098.90884° , 1520 m , August 20-24, 2005 , G. Tang ( HNU , CASENT9025826 ) ; 1♂ , Gaoligongshan, Shibali Forest Station , in moist stream gully, N27.16894° , E98.77205° , 2650 m , May 9, 2004 , C. Griswold & H.M. Yan ( HNU , CASENT9019903 ) ; 2♂ , Gaoligongshan, 7.41 km 315° WNW of Shibali, 36 km NNW of Fugong , beneath objects amidst dormant bamboo, along snowfield and avalanche debris, N27.20629° , E98.72001° , 3336 m , May 7, 2004 , C. Griswold & D. Kavanaugh ( CAS , CASENT9019946 ) ; 1♂ , Gaoligongshan, 6.18 km 280° W of Shibali , turning rocks in open meadow along stream, N27.18413° , E98.72024° , 3100 m , May 7, 2004 , C. Griswold & D. Kavanaugh ( HNU , CASENT9020705 ) ; 1♂ , Gaoligongshan, 7.41 km WNW of Shibali, 36 km NNW of Fugong , beneath objects amidst dormant bamboo, along snowfield and avalanche debris, N27.20629° , E98.72001° , 3336 m , May 8, 2004 , C. Griswold & D. Kavanaugh . ( HNU , CASENT9019947 ) ; 1♂ , Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 km above Shibali , N27.16784° , E98.77782° , 3059 m , May 4, 2004 , H.M. Yan , G.X. Peng ( CAS , CASENT9020580 ) ; 1♂ , Gaoligongshan, 10.1-11.5 road km from Shibali on Yaping Road , N27.20049° , E98.71354° - N27.20676° , E98.71763 , 3225-3290 m , night collecting along road, May 8, 2004 , C. Griswold & D. Kavanaugh ( HNU , CASENT9020032 ) . FIGURES 370-374. Draconarius pseudobrunneus Wang 2003, male (370-372) ( CAS , CASENT9020464) and female (373, 374) ( CAS , CASENT9019964) from Fugong County, Shibali Forest Station. 370. Habitus, dorsal; 371, 373. Eyes, fronto-dorsal; 372, 374. Labium, ventral ( L / W =1.00). FIGURES 375-386. Draconarius pseudobrunneus Wang 2003, female epigynum. 375-376. From Fugong County, Shibali ( CAS , CASENT9020627); 377-378. From Gongshan County, Cikai Township ( CAS , CASENT9025843); 379-380. From Fugong County, Shibali Forest Station ( HNU , CASENT9019963); 381-382. From Fugong County, Shibali Forest Station ( HNU , CASENT9019905); 383-384. From Fugong County, Maxidi ( CAS , CASENT9020583); 385-386. From Lushui County, Pianma Township, Yaojiaping Forest Station ( CAS , CASENT9022057). 375, 377, 379, 381, 383, 385. Ventral; 376, 378, 380, 382, 384, 386. Dorsal. FIGURES 387-398. Draconarius pseudobrunneus Wang 2003, female epigynum. 387-388. From Gongshan County, Cikai Township ( HNU , CASENT9025834); 389-390. From Gongshan County, Cikai Township ( CAS , CASENT9025841); 391-392. From Fugong County, Shibali Forest Station ( HNU , CASENT9019964); 393-394. From Fugong County, Shibali Forest Station ( HNU , CASENT9020459); 395-396. Label missing ( CAS , CASENT9020845); 397-398. From Lushui County, Feng Xue Yakou ( HNU , CASENT9022263). 387, 389, 391, 393, 395, 397. Ventral; 388, 390, 392, 394, 396, 398. Dorsal. FIGURES 399-406. Draconarius pseudobrunneus Wang 2003, female epigynum. 399-400. From Lushui County, Pianma Township, Feng Xue Yakou ( HNU , CASENT9022040); 401-404. From Gongshan County, Cikai Township ( CAS , CASENT9025836); 405-406. From Lushui County, Pianma Township, Feng Xue Yakou ( CAS , CASENT9020073). 399, 401, 403, 405. Ventral; 400, 402, 404, 406. Dorsal. Diagnosis: This species is similar to D. capitulatus Wang 2003 in having similar spermathecae, absence of epigynal teeth, and anteriorly extending conductor, and prolaterally originated embolus. The female of this species has the atrial ridges extending laterally, ending posteriorly close to the epigastric furrow, forming a round-shaped median area (Figs 407-408), whereas the atrial ridges of D. capitulatus extend relatively medially, ending posteriorly anterior to the epigastric furrow, and forming a triangular-shaped median area (Fig. 44). The male can be recognized by the slender patellar apophysis, the slightly bifurcate, smoothly curving conductor, and the absence of a median apophysis (Figs 409-410). Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 411-414, 418-424). Male (male of CASENT9025826 measured). The male is here described for the first time. Large sized coelotine, total length 12.04 (Fig. 425). Dorsal shield of prosoma 5.41 long, 3.63 wide; opisthosoma 6.63 long, 4.89 wide. AME smallest, half the size of ALE; ALE largest, PLE slightly smaller than ALE, PME slightly smaller than PLE ( AME 0.11, ALE 0.19, PME 0.16, PLE 0.18); AME separated from each other by their diameter, from PLE by slightly less than AME diameter; PME separated from each other by slightly less than their diameter, from PLE by slightly more than PLE diameter (AME-AME 0.11, AME-ALE 0.09, PME-PME 0.13, PME-PLE 0.20, AME-PME 0.14) (Fig. 426). Labium length and width subequal ( L / W =1.0) (Fig. 427). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with patellar apophysis long, slender; RTA short, less than or about half of tibial length, with distinctly protruding distal end; lateral tibial apophysis large; cymbial furrow small, less than half of cymbial length; conductor moderately long, anteriorly extending, with a slightly bifurcate apex, a small basal lamella, and a dorsal apophysis; median apophysis absent; embolus short, filiform, prolateral in origin (Figs 409-410, 415-417). Distribution: China (Yunnan: Fugong, Gongshan) (Fig. 547).