New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)AuthorPerrichot, VincentAuthorOrtega-Blanco, JaimeAuthorMcKellar, Ryan C.AuthorDelclos, XavierAuthorAzar, DanyAuthorNel, AndreAuthorTafforeau, PaulAuthorEngel, Michael S.textZooKeys2011130421453http://dx.doi.org/10.3897/zookeys.130.1453journal articlehttp://dx.doi.org/10.3897/zookeys.130.14531313-2970-130-421
Family
Maimetshidae Rasnitsyn, 1975Maimetshidae
Rasnitsyn, 1975: 73. Type genus:
Maimetsha
Rasnitsyn, 1975.
Rasnitsyn 1988
: 124;
Ronquist et al. 1999
: 33;
Rasnitsyn and Brothers 2009
: 192;
Perrichot 2009
: 2;
Ortega-Blanco et al. 2010
: 266;
Vilhelmsen et al. 2010a
: 674.
Dinapsinae
[
Megalyridae
], partim:
Shaw 1988
: 107.
Dinapsini
[
Megalyridae
], partim:
Shaw 1990
: 578.
Diagnosis.
Modified from
Rasnitsyn and Brothers (2009)
: Head hypognathous, without ocular carina, without subantennal groove accommodating antennal base; vertex with or without longitudinal median sulcus; antenna filiform, with scape short to moderate, never elongate, flagellomeres variable in number (14-19), most often 16, without apparent sexual dimorphism; mandibles often asymmetrical, with 3-4 teeth. Pronotum short medially; mesoscutum with notauli and median sulcus; axillae meeting anterior to mesoscutellum or separated by scutoscutellar sulcus; propodeum areolate. Forewing with costal space moderate to wide; C and pterostigma present; basal sections of Rs and M subequal in length, not continuously aligned (i.e., not forming smooth basal vein); cell [2R1] (i.e., marginal cell) closed, wide (not triangular), moderately short to moderately long; cells [1Rs] and/or [2Rs] closed; cell [1M] closed, small, distant from [1Rs]; cell [2M] open or delimited by spectral or nebulous 2m-cu; 1cu-a antefurcal. Hind wing with no posterobasal lobe; 4-5 distal hamuli; basal cell [R] closed; free apex of Rs present, short to long; free apices of M and Cu short or absent, that of A absent. Legs with trochantelli; tibial spur formula 1-2-2; tarsi pentamerous; tarsal plantulae present in some females; pretarsal claws with preapical tooth. Metasoma rather short, compact, not much sculptured, attached low on propodeum, first segment longest [articulatory ring referred to by
Rasnitsyn and Brothers (2009)
is not
evident
and apparently an incorrect interpretation], apical sternum of female elongate, nearly reaching metasomal apex; ovipositor external but not very long, sheaths at most as long as metasoma, not fitting tightly to ovipositor (often preserved detached).
Included genera.Andyrossia
Rasnitsyn and Jarzembowski, 2000 (Wealden, England, Barremian; a replacement name for
Arossia
Rasnitsyn and Jarzembowski in
Rasnitsyn et al. 1998
);
Ahiromaimetsha
Perrichot, Azar, Nel, and Engel gen n. (Lebanese amber, Neocomian);
Iberomaimetsha
Ortega-Blanco, Perrichot, and Engel gen. n. (Spanish amber, Lower Albian);
Guyotemaimetsha
Perrichot et al., 2004a (French amber, latest Albian/earliest Cenomanian);
Afrapia
Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian);
Afromaimetsha
Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian);
Maimetshorapia
Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian);
Maimetsha
Rasnitsyn, 1975 (Taimyr amber, Santonian); and
Ahstemiam
McKellar and Engel gen. n. (Canadian amber, Campanian). Table 1 summarizes the species diversity of the family.
Table 1. Known records for
Maimetshidae
s.l. (A = amber inclusion; C = compression fossil).