Parasopubia hofmannii Pradeep & Pramod and Parasopubia hofmannii var. albiflora Pradeep & Pramod (Orobanchaceae), two new taxa from India
Author
Pradeep, Ayilliath K.
Department of Botany, University of Calicut, Kerala- 673 635, India.
akpradeep1@rediffmail.com
Author
Pramod, Choorakkay
Department of Botany, Govt. Brennen College, Darmadam, Thalassery, Kerala- 670 106, India.
text
Candollea
2013
2013-07-01
68
1
115
122
journal article
3439
10.15553/c2013v681a16
53a3a5b6-86cb-462e-9e16-1ade7ac70634
2235-3658
5706265
Parasopubia hofmannii
var.
albiflora
Pradeep & Pramod,
var. nova
(
Fig. 4
,
5
).
Typus
:
INDIA
.
Kerala
:
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
1.VIII.2010
,
Pramod
CU 126522
(holo-:
G
; iso-:
MH
,
CALI
).
Closely allied to
P. hofmannii
var.
hofmannii
,
but can easily be distinguished by its elegant white corolla, plant parts without purplish tinge and seeds with distinctly polygonal testacellswithsecondary cellsand slightlyrugose walls.
Erect
annual herb, to
50 cm
high.
Stem
stiff, terete at base, tetragonous and sulcate above, upwards with erecto-pendent branches, diameter
3 mm
at base, glabrous, green; internodes
5-20 mm
long.
Leaves
opposite,
1-6 cm
long, lower leaves 3-linear segmented, upper leaves entire; margins entire, midrib depressed, strigose on margins, glabrous on both surfaces; segments
0.5 mm
broad, filiform, green, terminal segments as long as lateral segments or longer; leaves reduced towards apex.
Flowers
axillary, solitary, 1-1.5 ×
0.8-1.1 cm
; pedicels up to
3 mm
long.
Bracteoles
2, linear/subulate, 0.5 ×
2-6 mm
, strigose on margins, glabrous on both surfaces, tip acute.
Calyx
6 mm
long, green; lobes 5, unequal, tube
2 mm
long, 10-ribbed, glabrous; lobes linear-lanceolate,
2-4 mm
long, acute, margins strigose, glabrous on both surfaces, green, persistent in fruits.
Corolla
campanulate, 1-1.2 ×
0.7-1 cm
, white, corolla tube narrow (
1 mm
) up to
2 mm
of length of the tube, then expanding, lobes 5, suborbicular, 5 ×
4 mm
, minutely puberulous outside, glabrous within.
Stamens
4, unequal, filaments attached below the middle of the corolla tube, white, glabrous, 2 longer (
5 mm
), 2 shorter (
3 mm
), arched, attached above longer stamens.
Anthers
2, pendant from the apex of filament, all almost at same level, one perfect, oblong, shortly apiculate,
1.5-2 mm
long, white, dehiscence from base, cuspidate; sterile anther lanceolate,
1.5-2 mm
long, cuspidate.
Ovary
ellipsoid, 1.5 ×
1 mm
, green, glabrous, cells 2, placentation axile; ovules many.
Style
simple,
6 mm
long, hyaline, glabrous; stigma subglobose, hyaline, glabrous.
Capsules
obovoid, 4-5
3-3.5 mm
, green, much exceeding the calyx tube, emarginate at apex, apiculate with withered style, glabrous; fruiting calyx to
8 mm
long, lobes
5-6 mm
long, tube
2-3 mm
long, green.
Seeds
many, oblong,
0.75 mm
long, brownish, glabrous, testa cells polygonal, broad, secondary cells well developed with oblique walls, cell walls slightly rugose.
Phenology. –
Flowering and fruiting in June to October.
Specimina visa. –
INDIA
.
Kerala
:
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
11.IX.2009
,
Pramod
CU 123591
(
CALI
);
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
20.VIII.2010
,
Pramod
CU 126554
(
CALI
);
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
25.VII.2011
,
Pramod
CU 126690
(
CALI
);
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
1.VIII.2010
,
Pramod
CU 126507
(
CALI
);
Kannur
,
Madayippara
,
36 m
,
12º01.792’N
75º15.246’E
,
1.VIII.2010
,
Pramod
CU 126505
(
CALI
).
Tamil Nadu
:
Madurai
,
Thandigudi
,
1400 m
,
22.X.1977
,
M. Chandrabose
51625
(
MH
).
Taxonomical notes. –
Parasopubia hofmannii
var.
albiflora
is distinct in having elite white corolla and dark green plant parts against the purple corolla and purplish green plant parts of
var.
hofmannii
(
Fig. 2E
;
Fig. 5B
). The interpretation of the SEM images of seeds can also lend further support for the establishment of the new variety. In
var.
hofmannii
,
the testa cells are very narrow with smooth walls and without or ill-developed secondary ridges (
Fig. 3
C-D). In
var.
albiflora
, the testa cells are polygonal with slightly rugose walls and well developed secondary walls (
Fig. 3
E-F).