Berlese's Primitive Oribatid Mites Author van der Hammen, L. text Zoologische Verhandelingen 1959 40 1 93 http://www.repository.naturalis.nl/document/148866 journal article ORI111 0DC6B575-3CB3-41C1-A3EC-850520AE4487 Heterochthonius Berlese, 1910 Heterochthonius Berlese, 1910, pp. 221, 222. Berlese (1910) created Heterochthonius as a subgenus of Cosmochthonius ; he mentioned the simple dorsal hairs and the long tarsi as differential characters. Ewing (1917, p. 125) already regarded Heterochthonius as a separate genus, which opinion was established by Grandjean (1928) in his detailed description of H. gibbus , the type species.