Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Part 2
Author
Neubauer, Thomas A.
Author
Harzhauser, Mathias
Author
Kroh, Andreas
Author
Elisavet, Georgopoulou
Author
Mandic, Oleg
text
ZooKeys
2014
429
13
46
http://dx.doi.org/10.3897/zookeys.429.7420
journal article
http://dx.doi.org/10.3897/zookeys.429.7420
1313-2970-429-13
794E5F42F746425F996D5C6E64F89194
794E5F42F746425F996D5C6E64F89194
Taxon classification Animalia ORDO FAMILIA
Muellerpalia haszprunari
nom. n.
Valvata simplex
Fuchs, 1870 in
Fuchs 1870b
: 535, pl. 21, figs 4-6 [non
Valvata tricarinata
var. simplex Gould, 1841].
Valvata (Valvata) simplex simplex
Fuchs;
Wenz 1928a
: 2474 (cum syn.) [non
Gould 1841
].
Valvata simplex
Fuchs;
Strausz 1942
: 80 [non
Gould 1841
].
Hauffenia simplex
(Fuchs);
Schlickum 1978
: 247, pl. 18, fig. 3 [non
Gould 1841
].
Hauffenia simplex
(Fuchs 1870);
Harzhauser and Binder 2004
: 9 [non
Gould 1841
].
Etymology.
In honor of Gerhard Haszprunar (Bavarian State Collection of Zoology Munich and Ludwig Maximilians University Munich), who summarized all existing names of living and fossil valvatids in a comprehensive nomenclator (
Haszprunar 2014
).
Type
locality.
Tihany at Lake Balaton,
Veszprem
, Hungary.
Age.
Late Miocene (Late Pannonian, Transdanubian sensu
Sacchi and
Horvath
2002
;
Sztano
et al. 2013
).
Type material.
According to the inventory books of the Natural History Museum Vienna the material should be stored there, but despite great effort it could not be located.
Discussion.
This species is a primary homonym of the extant taxon
Valvata tricarinata
var. simplex Gould, 1841 (p. 226) from Massachusetts, USA. The American taxon was elevated to species level by
Fluck (1932)
. As the European species was combined with various genera since its first description and several subspecies have been described, a summary of its history is given below.
Already
Schlickum (1978)
considered
Valvata simplex
Fuchs, 1870 to belong to the
Hydrobiidae
and placed it in the genus
Hauffenia
, based on similarities of morphology and size. Recently,
Bandel (2010)
introduced the new genus
Muellerpalia
for
Valvata bicincta
Fuchs, 1870 in
Fuchs (1870b)
,
Valvata carinata
Fuchs, 1870 in
Fuchs (1870b)
,
Planorbis varians
Fuchs, 1870 in
Fuchs (1870a)
,
Valvata simplex
Fuchs, 1870 in
Fuchs (1870b)
, and two new species (see discussion in
Neubauer et al. 2014
for the rather confusing systematics applied in
Bandel 2010
). We follow Bandel and place the species within
Muellerpalia
.
The following subspecies have been introduced or ranked within
Valvata simplex
Fuchs, 1870:
1)
Valvata bicincta
Fuchs, 1870 [erroneously "bicinata" on p. 536; from captions and description there is no doubt about the correct name] from Tihany: It was considered a subspecies of
Valvata simplex
by
Lorenthey
(1906
, p. 166), what was followed by
Wenz (1928a
, p. 2475) and
Strausz (1942
, p. 36).
Bandel (2010
, p. 103) treated it as separate species and combined it with the new genus
Muellerpalia
. Current status:
Muellerpalia bicincta
.
2)
Valvata carinata
Fuchs, 1870 (p. 536) from Tihany: It was considered as subspecies of
Valvata simplex
by
Pană
et al. (1981)
and
Pană
(2003)
, but recombined with the new genus
Muellerpalia
by
Bandel (2010
, p. 104). It is, however, a primary homonym of
Valvata carinata
Sowerby, 1834 (replacement name is introduced below).
3)
Valvata simplex
oecsensis
Soos
, 1934 (p. 189) from
Oecs
:
Schlickum (1978
, p. 246) clearly separated this taxon from "
Hauffenia simplex
" and retained it in
Valvata
.
Wenz and Edlauer (1942
, p. 83) elevated it to species level, what was followed by
Papp (1953
, p. 109), and
Harzhauser and Binder (2004
, p. 10). In some of the mentioned publications the name was erroneously emended to "oecsensis"; the correct emendation following ICZN rules is "ocsensis", since it is not derived from a German expression (
ICZN 1999
, Article 32.5.2.1). Current status:
Valvata ocsensis
.
4)
Valvata octonaria
Brusina, 1902 (pl. 13) from Tihany: It was also ranked as subspecies of
Valvata simplex
by
Wenz (1928a
, p. 2476). Since it was not referred to by
Bandel (2010)
, its generic affiliation is uncertain. Current status (needs revision):
Muellerpalia haszprunari octonaria
.
5
)
Valvata simplex polycincta
Lorenthey
, 1906 (p. 167) from Tihany: It was synonymized with
Valvata simplex octonaria
by
Wenz (1928a
, p. 2476). Current status: junior synonym of
Muellerpalia haszprunari octonaria
.
6)
Valvata simplex unicincta
Lorenthey
, 1906 (p. 165) from Tihany (
Feherpart
): The status of this taxon is doubtful. It was not mentioned by
Wenz (1928a
) or
Bandel (2010)
. Given the similarity with simplex and bicincta stated by
Lorenthey
, it might fall into the intraspecific variability of either of these species. Current status (needs revision):
Muellerpalia haszprunari unicincta
.