The open-holed trapdoor spiders (Mygalomorphae: Anamidae: Namea) of Australia’s D’Aguilar Range: revealing an unexpected subtropical hotspot of rainforest diversity Author Rix, Michael G. 0000-0001-5086-3638 Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD 4101, Australia. & Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia. michael.rix@qm.qld.gov.au Author Wilson, Jeremy D. Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD 4101, Australia. & Division of Arachnology, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470 (C 1405 DJR), Buenos Aires, Argentina. Author Harvey, Mark S. Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia. & School of Animal Biology, The University of Western Australia, Crawley, WA 6009, Australia. text Zootaxa 2020 2020-10-15 4861 1 71 91 journal article 8280 10.11646/zootaxa.4861.1.5 38255d4a-0223-4819-9a6c-cf3e93c57446 1175-5326 4414567 44321429-80FA-45AC-90D6-E3E13C961BFC Namea gloriosa sp. nov. ( Figs 14a, b , 22–34 ) http://zoobank.org/?lsid= urn:lsid:zoobank.org:act: F2560FF4-4484-4A5D-A2EB-4D8D588749DD Namea cucurbita Raven, 1984: 22 (in part; cited paratype specimen QMB S799 from Mt Tenison Woods). Type material. AUSTRALIA : Queensland : male holotype , Mount Glorious , malaise trap , rainforest, 10 Au-gust–13 November 1983 , A. Hiller ( QMB S10269 ) . Paratype : 1 male , Tenison Woods Mountain [sic “Mt Tenison Woods”], pitfall trap , 26 June–18 November 1978 , G. & S. Monteith ( QMB S799 ) . Diagnosis. Males of Namea gloriosa can be distinguished from those of all other described congeners except N. cucurbita by the size of the isolated, proximal retroventral macroseta on the palpal tibia, which is unusually long (i.e. clearly longer than the length of the palpal bulb) ( Fig. 32 ). Males can be further distinguished from those of N. cucurbita by the shorter embolus ( Fig. 34 ; cf. Rix et al. 2020 , fig. 41) and the more slender profile of tibia I ( Fig. 31 ; cf. Rix et al. 2020 , fig. 38). Females are unknown. Description (male holotype ): Total length 19.9. Carapace 7.7 long, 6.1 wide. Abdomen 8.5 long, 4.4 wide. Carapace ( Fig. 22 ) dark chocolate-brown, covered with reflective downy setae ( Fig. 24 ); lateral margins with fringe of anteriorly curved, porrect black setae, longest posteriorly; fovea straight. Eye group ( Fig. 25 ) rectangular, twice as wide as long, PLE–PLE/ALE–ALE ratio 1.0; AME separated by less than their own diameter; PME separated by 4.1 X their own diameter; PME and PLE almost contiguous. Maxillae each with field of ca. 70 cuspules confined to heel and inner proximal corner ( Fig. 26 ); labium without cuspules. Abdomen ( Figs 23, 28 ) elongate-oval, dark brown with course pale beige-brown marbled pattern dorsally, and pale beige-brown ventrally with finer dark brown mottling; covered with short, fine setae. Legs ( Figs 22, 29–31 ) dark brown, with light scopulae on tarsi I–IV and distal half of metatarsi I–II; tibia I with 3 prodorsal, 2 proventral, 3 ventral and 1 retrolateral macrosetae; macroseta v1 large, reaching beyond ventro-distal margin of tibia I ( Fig. 31 ). Leg I: femur ca. 5.7 [damaged], patella 4.0, tibia 4.6, metatarsus 4.7, tarsus 3.1, total length ca. 22.1. Leg I femur–tar-sus/carapace length ratio ca. 2.9. Pedipalpal tibia ( Figs 32–34 ) 2.5 X longer than wide, with isolated, proximal retroventral macroseta (which is longer than length of palpal bulb), 1 retrodistal macroseta, 1 prodistal macroseta, and 2 proventral macrosetae. Cymbium ( Figs 32–34 ) setose, with distal scopula. Bulb ( Figs 32–34 ) sub-spherical, with long, whip-like, reflexed embolus arising from retroventral base of bulb. Etymology. The specific epithet is derived from the Latin gloriosus (adjective: ‘glorious’, ‘full of glory’), in reference to the type locality of this species. Distribution. Namea gloriosa is endemic to the D’Aguilar Range, where it is known only from rainforest at Mount Glorious and Tenison Woods Mountain ( Fig. 2 ). Remarks. This species is extremely rare, and currently known from only two male specimens. Nothing is known of its biology or life history, other than that the two known male specimens were collected wandering in search of females in winter or spring.